Boechera collinsii |
Boechera platysperma |
|
---|---|---|
collin's suncress, Collins' rockcress |
broad-seed rock-cress, flat-seed rock-cress, pioneer rock-cress |
|
Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, elevated on woody base above or near ground surface, 0.6–2.8(–3.5) dm, densely pubescent proximally, trichomes short-stalked, 2–5-rayed, 0.1–0.3 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade narrowly oblanceolate, 3–7(–10) mm wide, margins entire, ciliate near petiole base, trichomes (simple or spurred), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 2–4 (or 5)-rayed 0.1–0.3 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
3–7(–12), not concealing stem; blade auricles absent, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | (15–)30–90-flowered, usually unbranched. |
2–7-flowered, unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals creamy white to purplish, 4–6 × 1–2 mm, sparsely pubescent (often some trichomes abaxially); pollen ellipsoid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
ascending, straight, 3–13 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
ascending, not appressed to rachis, not secund, straight to slightly curved, edges undulate (not parallel), (2.5–)4–8.5 cm × 3–5.5 mm; valves glabrous; ovules 16–44 per ovary; style 0.05–1 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
uniseriate, 3–6(–8) × 2–3.5 mm; wing continuous, 0.8–2.5 mm wide. |
2n | = 14. |
= 14. |
Boechera collinsii |
Boechera platysperma |
|
Phenology | Flowering May–Jun. | Flowering Jun–Aug. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Rock outcrops and gravelly soil in dry pine forests and lodgepole-chaparral woodlands |
Elevation | 600-1400 m (2000-4600 ft) | 1600-3000 m (5200-9800 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
|
CA; OR
|
Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Previous authors have adopted broader circumscriptions of Boechera platysperma, including plants herein called B. covillei, B. elkoensis, and B. howellii. In our opinion, the differences between these taxa are sufficient to justify their treatment as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007, 2007b for detailed comparison). Reports of B. platysperma from Nevada (e.g., R. C. Rollins 1993; N. H. Holmgren 2005b) are based on plants of B. howellii or apomictic hybrids with other species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 397. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis platysperma, Arabis inamoena, Arabis oligantha |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (A. Gray) Al-Shehbaz: Novon 13: 388. (2003) |
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