Boechera collinsii |
Boechera pauciflora |
|
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collin's suncress, Collins' rockcress |
Columbia rockcress, few-flower rock-cress, fuzzy suncress, hairy-stem rock-cress, small-flower rockcress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; short- to long-lived; apomictic; caudex sometimes woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.4–)3–11.2 dm, densely pubescent proximally, trichomes simple, 0.6–1.5 mm, mixed with stalked, 2- (or 3-)rayed ones, 0.2–0.4 mm, sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade oblanceolate, 3–10 mm wide, margins usually dentate, rarely entire, sometimes ciliate, trichomes (simple or branched), to 1 mm, surfaces densely pubescent, trichomes stalked, 2–5-rayed, 0.3–0.6 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
(8–)14–60, often concealing stem proximally; blade auricles (1–)3–10 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | (15–)30–90-flowered, usually unbranched. |
17–60-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
divaricate at anthesis; sepals pubescent; petals lavender to whitish, 5–8 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
horizontal to divaricate-descending, usually straight, rarely slightly recurved, 4–13 mm, pubescent, trichomes spreading, 2- or 3-rayed. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
horizontal, divaricate-descending or widely pendent, not appressed to rachis, not secund, curved, edges parallel, 5.5–10.5 cm × 1.5–2.2 mm; valves glabrous; ovules 80–162 per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
uniseriate, 1.4–1.8 × 1–1.4 mm; wing continuous, 0.1–0.25 mm wide. |
2n | = 14. |
= 21. |
Boechera collinsii |
Boechera pauciflora |
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Phenology | Flowering May–Jun. | Flowering May–Jun. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Rocky soil in sagebrush areas, mountain shrub communities, edges of conifer forests |
Elevation | 600-1400 m (2000-4600 ft) | 600-2500 m (2000-8200 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
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CA; ID; MT; NV; OR; UT; WA; WY; BC
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Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera pauciflora is an apomictic species that arose through hybridization between B. retrofracta and B. sparsiflora. Specimens of B. pauciflora are commonly identified as Arabis holboellii var. pinetorum (= B. pinetorum), a superficially similar species restricted to the northern Sierra Nevada and southern Cascade Range (see M. D. Windham and I. A. 2007 for detailed comparison). Arabis elegans A. Nelson (1900), not Tineo & Lojacono (1886) is an illegitimate name, sometimes found in synonymy with Boechera pauciflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 393. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Sisymbrium pauciflorum, Arabis arcuata var. subvillosa, Arabis columbiana, Arabis perelegans, Arabis sparsiflora var. columbiana, Arabis sparsiflora var. subvillosa, B. sparsiflora var. subvillosa |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 268. (2007) |
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