Boechera collinsii |
Boechera lemmonii |
|
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collin's suncress, Collins' rockcress |
Lemmon's rock-cress, Lemmon's suncress, wind-blown suncress |
|
Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; long-lived; (somewhat cespitose); sexual or apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.5–2(–2.5) dm, glabrous or sparsely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.1–0.2 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade oblanceolate to obovate, 1.5–5 mm wide, margins usually entire, rarely slightly dentate, ciliate along petiole, surfaces densely to sparsely pubescent, trichomes short-stalked, 3–9-rayed, 0.1–0.2 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
2–8(–12), not concealing stem; blade auricles absent or 0.1–0.5 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | (15–)30–90-flowered, usually unbranched. |
3–12(–17)-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous or sparsely pubescent; petals purple to lavender, 3.5–6 × 1–1.5 mm; pollen ellipsoid or spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
divaricate-ascending to slightly descending, usually slightly recurved, rarely straight, 2–6 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
divaricate-ascending to slightly descending, not appressed to rachis, secund, straight or curved, edges parallel, (1.6–) 2–4.4 cm × 1.6–2.3 mm; valves glabrous; ovules 28 40(–44) per ovary; style 0.1–0.2 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
uniseriate, 1.3–2 × 1–1.5 mm; wing continuous, 0.1–0.5 mm wide. |
2n | = 14. |
= 14. |
Boechera collinsii |
Boechera lemmonii |
|
Phenology | Flowering May–Jun. | Flowering Jun–Aug. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Cliffs, talus slopes, and gravelly soil in alpine and subalpine habitats |
Elevation | 600-1400 m (2000-4600 ft) | 2100-4400 m (6900-14400 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
|
AK; CA; CO; ID; MT; NV; OR; UT; WA; WY; AB; BC; YT
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Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera lemmonii is easily recognized by its combination of secund fruits, mat-forming habit, purplish sepals, and obovate-oblanceolate cauline leaves. Both sexual and apomictic collections are known; further study is needed to determine whether they truly are conspecific. The taxa traditionally treated as Arabis (Boechera) lemmonii vars. depauperata, drepanoloba, and paddoensis are apomictic hybrids here recognized as distinct species (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 388. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis lemmonii, Arabis bracteolata, Arabis canescens var. latifolia, Arabis codyi, Arabis egglestonii, Arabis kennedyi, Arabis latifolia, Arabis oreocallis, Arabis polyclada, Arabis semisepulta |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (S. Watson) W. A. Weber: Phytologia 51: 370. (1982) |
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