Boechera collinsii |
Boechera horizontalis |
|
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collin's suncress, Collins' rockcress |
Crater Lake rock-cress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; long-lived; (cespitose); apomictic; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, near ground surface or somewhat elevated on woody base, or 1–3.5 dm, sparsely pubescent proximally, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade narrowly oblanceolate, 1–5 mm wide, margins entire, ciliate near petiole base, trichomes (simple), to 0.4 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.1–0.3 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
3–13, often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | (15–)30–90-flowered, usually unbranched. |
5–32-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals lavender to purple, 5–6 × 1.5–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
horizontal to descending, straight or slightly curved downward, 4–11 mm, glabrous. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
horizontal or descending, not appressed to rachis, secund, straight, edges slightly undulate (not parallel), 2–4 cm × 2–3 mm; valves glabrous; ovules 40–54 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
uniseriate, 2–2.5 × 1.7–2 mm; wing continuous, 0.5–1 mm wide. |
2n | = 14. |
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Boechera collinsii |
Boechera horizontalis |
|
Phenology | Flowering May–Jun. | Flowering Jul–Aug. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Dry pumice slopes |
Elevation | 600-1400 m (2000-4600 ft) | |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
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OR |
Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera horizontalis is an apomictic species that arose through hybridization between B. lemmonii and B. suffrutescens (M. D. Windham and I. A. Al-Shehbaz 2007). Boechera horizontalis is known only from the vicinity of Crater Lake in south-central Oregon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 384. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis horizontalis, Arabis suffrutescens var. horizontalis |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) |
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