Boechera collinsii |
Boechera harrisonii |
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collin's suncress, Collins' rockcress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; long-lived; (some-what cespitose); apomictic; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 3–7 per caudex branch, arising from margin of rosette near ground surface, 0.5–2.5 dm, usually sparsely pubescent proximally, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.06–0.2 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade narrowly oblanceolate, 2–4 mm wide, margins entire or somewhat dentate, ciliate near petiole base, trichomes (simple), to 0.5 mm, surfaces sparsely pubescent, trichomes short-stalked, 2–5-rayed, 0.1–0.25 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
3–7, not concealing stem; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves glabrous. |
Racemes | (15–)30–90-flowered, usually unbranched. |
5–12-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender, 5–7.5 × 0.8–1.5 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
divaricate-ascending, straight, 8–12 mm, glabrous. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
usually divaricate-ascending, rarely nearly horizontal, not appressed to rachis, not secund, curved to straight, edges parallel, 3–4.7 cm × 1–1.5 mm; valves glabrous; ovules 64–86 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
uniseriate, 1.1–1.3 × 0.8–1.1 mm; wing continuous, 0.08–0.1 mm wide. |
2n | = 14. |
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Boechera collinsii |
Boechera harrisonii |
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Phenology | Flowering May–Jun. | Flowering May–Jun. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Limestone and quartzite cliffs |
Elevation | 600-1400 m (2000-4600 ft) | 1500-1600 m (4900-5200 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
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UT |
Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera harrisonii is an apomictic species that arose through hybridization between B. microphylla and B. perennans (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). Boechera harrisonii is known only from Utah and Wasatch counties in north-central Utah. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 383. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis harrisonii, B. microphylla var. harrisonii |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (S. L. Welsh) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 266. (2007) |
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