Boechera collinsii |
Boechera formosa |
|
---|---|---|
collin's suncress, Collins' rockcress |
beautiful rockcress |
|
Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex branch, arising from center of leaf tuft, above ground surface on woody base, 2–5.5 dm, densely pubescent proximally, trichomes short-stalked, 4–7-rayed, 0.1–0.4 mm, similarly pubescent distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade linear to linear-oblanceolate, 2–3(–4) mm wide, margins entire, sometimes ciliate near petiole base, trichomes (simple or 2-rayed), to 0.8 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
7–18, concealing stem proximally; blade auricles absent, surfaces of distalmost leaves densely pubescent. |
Racemes | (15–)30–90-flowered, usually unbranched. |
6–26-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
divaricate at anthesis; sepals pubescent; petals white to pale lavender, 8–18 × 2.5–4.4(–5.5) mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
horizontal to descending, usually recurved, 5–10(–20) mm, pubescent, trichomes appressed, branched. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
divaricate-descending to reflexed, usually not appressed to rachis, rarely somewhat secund, straight, edges parallel, 4.3–7 cm × 1.6–3 mm; valves pubescent throughout; ovules 72–98 per ovary; style ca. 0.1 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
biseriate, 1.2–1.6 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 14. |
= 14. |
Boechera collinsii |
Boechera formosa |
|
Phenology | Flowering May–Jun. | Flowering Apr–Jun. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Rocky slopes and sandy soil in blackbrush, sagebrush, and pinyon-juniper communities |
Elevation | 600-1400 m (2000-4600 ft) | 1300-1900 m (4300-6200 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
|
AZ; CO; NM; UT; WY
|
Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Boechera formosa is usually subsumed within Arabis (Boechera) pulchra, the two species are easily separated. Boechera formosa has white to lavender (versus mostly purple) petals, fruiting pedicels that are not abruptly recurved at the base, narrower (1.6–3 versus 2.5–4 mm wide), mostly non-appressed fruits, and narrower (1–1.2 versus 1.5–2.2 mm wide) seeds. The two taxa are separated by more than 500 km, with B. formosa found only on the Colorado Plateau and B. pulchra, in the strict sense, restricted to southern California, western Nevada, and northwestern Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 379. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis formosa, Arabis pulchra var. pallens, B. pulchra subsp. pallens, B. pulchra var. pallens |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 68. (2006) |
Web links |