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collin's suncress, Collins' rockcress

Fendler's rockcress

Habit Perennials; usually short-lived; sexual; caudex not woody. Perennials; short- to long-lived; sexual; caudex usually not woody (sometimes with persistent, crowded leaf bases).
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally.

1–7 per caudex branch, arising from center of rosette near ground surface, or arising laterally proximal to sterile shoots, 1.5–5.5(–8) dm, densely pubescent proximally, trichomes simple and long-stalked, 2-rayed, 0.2–0.9 mm, glabrous distally.

Basal leaves

blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm.

blade broadly oblanceolate, 5–15(–20) mm wide, margins dentate, ciliate, trichomes (simple), to 1.2 mm, surfaces glabrous or pubescent, trichomes simple and long-stalked, usually 2-rayed, rarely some 3-rayed, 0.2–0.6 mm.

Cauline leaves

(8–)15–40, concealing stem proximally;

blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent.

8–25, often concealing stem proximally;

blade auricles 0.8–3 mm, surfaces of distalmost leaves often glabrous.

Racemes

(15–)30–90-flowered, usually unbranched.

6–40(–74)-flowered, usually unbranched.

Flowers

descending to pendent at anthesis;

sepals pubescent;

petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous;

pollen ellipsoid.

ascending at anthesis;

sepals sparsely hirsute;

petals usually lavender, rarely white, 5–9 × 1–2 mm;

pollen ellipsoid.

Fruiting pedicels

reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple.

horizontal to divaricate-ascending, curved or angled downward, 9–18(–23) mm, usually glabrous, rarely sparsely pubescent, trichomes spreading, simple.

Fruits

strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm;

valves glabrous;

ovules 70–126 per ovary;

style 0.2–0.5 mm.

widely pendent, not appressed to rachis, not secund, curved to nearly straight, edges parallel, 3–5.8 cm × 1.5–2 mm;

valves glabrous;

ovules 98–128 per ovary;

style 0.2–0.5 mm.

Seeds

uniseriate, 1–1.4 × 0.8–1 mm;

wing continuous (rarely absent), to 0.1 mm wide.

biseriate, 0.9–1.2 × 0.6–0.8 mm;

wing distal (sometimes absent), 0.05–0.15 mm wide.

2n

= 14.

= 14.

Boechera collinsii

Boechera fendleri

Phenology Flowering May–Jun. Flowering Apr–Jul.
Habitat Rocky and gravelly hillsides, prairies, open woods, floodplains Rocky slopes in pine forests, pinyon-juniper woodlands, scrub oak
Elevation 600-1400 m (2000-4600 ft) 1800-2800 m (5900-9200 ft)
Distribution
from FNA
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; NV; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

As circumscribed by R. C. Rollins (1993), Boechera fendleri included distinctive elements, segregated here as B. porphyrea, B. spatifolia, and B. texana (see M. D. Windham and I. A. Al-Shehbaz 2006, 2007 for detailed comparison). Boechera fendleri in the strict sense is a sexual diploid extending from the Four Corners region through northern Arizona to southern Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 370. FNA vol. 7, p. 378.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii Arabis holboellii var. fendleri, Arabis fendleri
Name authority (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) (S. Watson) W. A. Weber: Phytologia 51: 370. (1982)
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