Boechera collinsii |
Boechera duchesnensis |
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collin's suncress, Collins' rockcress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody. | Perennials; usually long-lived; apomictic; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from near ground surface, 1.5–4.5 dm, densely pubescent proximally, trichomes short-stalked, 2–7-rayed, 0.2–0.5 mm (sometimes mixed with some larger trichomes proximally), sparsely pubescent distally. |
Basal leaves | blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm. |
blade linear-oblanceolate, 2–5 mm wide, margins entire or denticulate, ciliate at petiole base, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.15–0.4 mm. |
Cauline leaves | (8–)15–40, concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent. |
3–8, not concealing stem; blade auricles 0.7–2 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | (15–)30–90-flowered, usually unbranched. |
11–22-flowered, usually unbranched. |
Flowers | descending to pendent at anthesis; sepals pubescent; petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals whitish to pale lavender, 7–10 × 1.5–3 mm, glabrous; pollen spheroid. |
Fruiting pedicels | reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple. |
horizontal, straight or slightly curved, 7–14 mm, pubescent, trichomes appressed, branched. |
Fruits | strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm; valves glabrous; ovules 70–126 per ovary; style 0.2–0.5 mm. |
horizontal, not appressed to rachis, not secund, usually straight, edges parallel, 3.5–5 cm × 1.7–2 mm; valves glabrous proximally, pubescent distally; ovules 72–92 per ovary; style 0.1–0.3 mm. |
Seeds | uniseriate, 1–1.4 × 0.8–1 mm; wing continuous (rarely absent), to 0.1 mm wide. |
sub-biseriate, 1.1–1.3 × 0.7–1 mm; wing continuous, ca. 0.1 mm wide. |
2n | = 14. |
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Boechera collinsii |
Boechera duchesnensis |
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Phenology | Flowering May–Jun. | Flowering May. |
Habitat | Rocky and gravelly hillsides, prairies, open woods, floodplains | Sandy soil on rocky slopes, mostly in pinyon-juniper woodlands |
Elevation | 600-1400 m (2000-4600 ft) | 1300-1800 m (4300-5900 ft) |
Distribution |
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
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CO; NM; UT |
Discussion | Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological and isozyme analyses indicate that Boechera duchesnensis is an apomictic species that arose through hybridization between B. formosa and B. pallidifolia (M. D. Windham and Allphin, unpubl.). It is easily distinguished from those species by having fruits that are glabrous proximally and sparsely pubescent distally. It shares this distinctive fruit pubescence with B. xylopoda, an apomictic hybrid between B. perennans and B. pulchra (see Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 370. | FNA vol. 7, p. 375. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii | Arabis pulchra var. duchesnensis, B. pulchra var. duchesnensis |
Name authority | (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) | (Rollins) Windham: Harvard Pap. Bot. 12: 239. (2007) |
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