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collin's suncress, Collins' rockcress

Four Corners rockcress

Habit Perennials; usually short-lived; sexual; caudex not woody. Biennials or perennials; short-lived; apomictic; caudex present or absent.
Stems

usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)2.5–8 dm, densely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 1 mm, sparsely pubescent or glabrous distally.

usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–5 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.2–0.5 mm, glabrous or sparsely pubescent distally.

Basal leaves

blade oblanceolate, 1–7 mm wide, margins entire or dentate, ciliate along petiole, trichomes (simple), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 5–8-rayed, 0.15–0.4 mm.

blade oblanceolate, 2–10 mm wide, margins usually dentate, sometimes ciliate near petiole base, trichomes (usually spurred), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.4 mm.

Cauline leaves

(8–)15–40, concealing stem proximally;

blade auricles 1–3 mm, surfaces of distalmost leaves usually sparsely pubescent.

15–36, sometimes concealing stem proximally;

blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent.

Racemes

(15–)30–90-flowered, usually unbranched.

20–55-flowered, usually unbranched.

Flowers

descending to pendent at anthesis;

sepals pubescent;

petals usually white, rarely lavender, 4–6 × 0.8–1.5 mm, glabrous;

pollen ellipsoid.

divaricate-descending at anthesis;

sepals pubescent;

petals pale lavender, 5–8.5 × 1–2 mm, glabrous;

pollen spheroid.

Fruiting pedicels

reflexed, abruptly recurved at base, otherwise straight, 4–13 mm, glabrous or sparsely pubescent, trichomes usually simple.

reflexed to divaricate-descending, usually curved downward, 8–14 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched.

Fruits

strongly reflexed, usually appressed to rachis, rarely somewhat secund, straight, edges parallel, 3.5–6 cm × 0.9–1.5 mm;

valves glabrous;

ovules 70–126 per ovary;

style 0.2–0.5 mm.

reflexed to pendent, rarely appressed to rachis, not secund, straight or slightly curved, edges parallel, 4–6 cm × 1–2 mm;

valves glabrous;

ovules 100–128 per ovary;

style 0.05–0.5 mm.

Seeds

uniseriate, 1–1.4 × 0.8–1 mm;

wing continuous (rarely absent), to 0.1 mm wide.

biseriate to sub-biseriate, mature seeds not seen.

2n

= 14.

Boechera collinsii

Boechera consanguinea

Phenology Flowering May–Jun. Flowering May.
Habitat Rocky and gravelly hillsides, prairies, open woods, floodplains Rocky slopes and sandy soil in ponderosa pine, pinyon-juniper, and sagebrush communities
Elevation 600-1400 m (2000-4600 ft) 1900-2500 m (6200-8200 ft)
Distribution
from FNA
ID; MN; MT; ND; NE; SD; WY; AB; MB; QC; SK
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
Discussion

Though often treated as a variety of Arabis (Boechera) holboellii (e.g., R. C. Rollins 1993; G. A. Mulligan 1996), B. collinsii is easily distinguished from that species by having simple and 2-rayed (versus 4–8-rayed) trichomes proximally on stems, and narrower (0.9–1.5 versus 2–2.5 mm) usually non-secund fruits. The two taxa also show allopatric distributions, with B. collinsii found on the North American continent (usually east of the Rocky Mountains) and B. holboellii apparently confined to Greenland.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Morphological evidence suggests that Boechera consanguinea is an apomictic species that arose through hybridization between B. fendleri and B. retrofracta. It is most similar to B. goodrichii, another apomictic hybrid involving B. retrofracta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 370. FNA vol. 7, p. 371.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Arabis collinsii, Arabis holboellii var. collinsii, Arabis retrofracta var. collinsii Arabis consanguinea, Arabis holboellii var. consanguinea
Name authority (Fernald) Á. Löve & D. Löve: Taxon 31: 125. (1982) (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 261. (2007)
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