Boechera breweri |
Boechera inyoensis |
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Brewer's rock-cress |
Inyo rockcress |
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Habit | Perennials; long-lived; sexual; caudex woody. | Perennials; short-lived; apomictic; caudex usually not woody. | ||||
Stems | usually 1 per caudex branch, arising from center of rosette, somewhat elevated on woody base or from ground surface, 0.6–3.5(–4.5) dm, densely pubescent proximally, trichomes long-stalked and simple, 2-rayed (sometimes all simple), 0.4–1 mm, sparsely pubescent or glabrous distally. |
usually 1 per caudex, arising from center of rosette near ground surface, (1–)2.5–6.5 dm, densely pubescent proximally, trichomes short-stalked, 7–12-rayed, 0.1–0.2 mm, densely to sparsely pubescent distally. |
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Basal leaves | blade oblanceolate, 3–7(–11) mm wide, margins entire or dentate, ciliate proximally, trichomes to 1 mm, surfaces densely pubescent, trichomes long-stalked, 2–4-rayed, 0.4–0.8 mm. |
blade oblanceolate, 1–4(–8) mm wide, margins entire, ciliate along petiole and proximally, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 3–10-rayed, 0.2–0.7 mm. |
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Cauline leaves | 5–15(–28), often concealing stem proximally; blade auricles 0.5–3(–5) mm, surfaces of distalmost leaves pubescent. |
(7–)12–35, concealing stem proximally; blade auricles 0.5–2 mm, surfaces of distalmost leaves densely pubescent. |
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Racemes | 7–20(–30)-flowered, usually unbranched. |
10–65-flowered, usually unbranched. |
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Flowers | ascending at anthesis; sepals pubescent; petals usually purple (rarely lavender), 7–12 × 2–4 mm, glabrous; pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals lavender to purplish, 5–8 × 1.2–2 mm, glabrous; pollen spheroid. |
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Fruiting pedicels | ascending to divaricate-ascending, straight, 3–25 mm, pubescent or glabrous, trichomes spreading, simple and 2-rayed. |
divaricate-ascending to horizontal, straight or slightly curved downward, 5–15 mm, sparsely pubescent, trichomes appressed, branched. |
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Fruits | ascending to divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.5–10 cm × 1.5–2.2 mm; valves usually glabrous, rarely sparsely pubescent; ovules 48–96 per ovary; style 0.05–0.3 mm. |
divaricate-ascending to widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 3.7–6.5 cm × 1.5–2.2 mm; valves glabrous; ovules 74–134 per ovary; style 0.05–0.2 mm. |
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Seeds | uniseriate, 1.2–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide distally. |
usually sub-biseriate, rarely uniseriate, 1.7–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
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2n | = 21. |
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Boechera breweri |
Boechera inyoensis |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Limestone and volcanic rock outcrops and clay soils in desert scrub and pinyon-juniper woodlands | |||||
Elevation | 1400-2400 m (4600-7900 ft) | |||||
Distribution |
CA; OR
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CA; NV; UT
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Discussion | Subspecies 2 (2 in the flora). Relatively few other sexual diploid species of Boechera co-occur with B. breweri, and the only currently recognized hybrid involving this taxon is B. rubicundula (= B. arcuata × B. breweri; M. D. Windham and I. A. Al-Shehbaz 2007). Boechera breweri appears to be related to B. sparsiflora, and some specimens of subsp. shastaensis can be difficult to separate from that species. The two subspecies appear to intergrade on the western side of the Sacramento Valley from Tehama to Solano counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Some plants here assigned to Boechera inyoensis were treated by R. C. Rollins (1993) and N. H. Holmgren (2005b) as Arabis (or B.) beckwithii. Closer examination of the type specimens of A. beckwithii reveals that they are conspecific with B. puberula, a sexual diploid widespread in the Great Basin of western North America. Boechera inyoensis is an apomictic triploid that clearly contains at least one genome derived from B. shockleyi (M. D. Windham and I. A. Al-Shehbaz 2007b); the other parent has not been determined. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 367. | FNA vol. 7, p. 385. | ||||
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Arabis breweri, Arabis breweri var. figularis, Arabis epilobioides | Arabis inyoensis, Arabis holboellii var. derensis, B. selbyi var. inyoensis | ||||
Name authority | (S. Watson) Al-Shehbaz: Novon 13: 384. (2003) | (Rollins) Al-Shehbaz: Novon 13: 386. (2003) | ||||
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