Boechera atrorubens |
Boechera divaricarpa |
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black-flower rockcress, dark-red-flower rockcress, purple sickle-pod rock-cress, sicklepod rockcress |
spreading-pod rock-cress, uplifting suncress |
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Habit | Perennials; usually short-lived; sexual; caudex not woody. | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette near ground surface, 0.8–6 dm, sparsely pubescent proximally, trichomes short-stalked and simple, (scattered), 2-rayed, 0.1–0.15 mm, glabrous distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1.5–)3–9 dm, glabrous or pubescent proximally, trichomes sessile, 2–4-rayed (some simple), to 0.7 mm, glabrous distally. |
Basal leaves | blade oblanceolate, 4–10 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 3- or 4-rayed, 0.1–0.2 mm. |
blade oblanceolate, 2–10 mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes (simple), to 0.8 mm, surfaces sparsely to densely pubescent, trichomes sessile, 2–6-rayed, 0.1–0.4 mm. |
Cauline leaves | 7–25, concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous. |
(10–)15–56, concealing stem proximally; blade auricles 1–5 mm, surfaces of distalmost leaves glabrous. |
Racemes | 6–20-flowered, usually unbranched. |
12–40(–65)-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals dark reddish purple (drying indigo), 7–9 × 1.5–2 mm, glabrous; pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous or with scattered trichomes; petals usually purple, rarely lavender, 6–9 × 1.5–3 mm, glabrous; pollen spheroid. |
Fruiting pedicels | ascending, usually straight, 5–10 mm, glabrous or pubescent, trichomes spreading, simple. |
divaricate-ascending to horizontal, straight, 5–12 mm, glabrous. |
Fruits | ascending, not appressed to rachis, not secund, curved or straight, edges parallel, (4–)6–12 cm × 1.7–2.2 mm; valves glabrous; ovules 80–100 per ovary; style 0.2–0.5 mm. |
divaricate-ascending to horizontal, not appressed to rachis, not secund, straight, edges parallel, (4.5–)5.5–11 cm × 1.7–2.5 mm; valves glabrous; ovules 114–142 per ovary; style 0.05–0.2 mm. |
Seeds | uniseriate, 1.2–1.7 × 0.9–1.2 mm; wing lateral and distal, 0.1–0.2 mm wide. |
uniseriate to sub-biseriate, 1.4–2 × 1–1.5 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 21. |
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Boechera atrorubens |
Boechera divaricarpa |
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Phenology | Flowering Apr–May. | Flowering May–Jul. |
Habitat | Rocky summits and sandy loam on sagebrush slopes | Rock outcrops, talus slopes and gravelly hillsides in sagebrush, mountain shrub, and open conifer forests |
Elevation | ca. 600 m (ca. 2000 ft) | 900-2500 m (3000-8200 ft) |
Distribution |
OR; WA
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CA; ID; MT; NV; UT; WA; WY
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Discussion | Boechera atrorubens is often treated as a variety of B. sparsiflora (e.g., R. C. Rollins 1993), it is readily separated from that species by having proximal stems sparsely (versus densely) pubescent with much smaller (0.15 versus 1.5 mm) trichomes. The two taxa rarely grow in proximity and, in areas where they are sympatric, B. atrorubens is further distinguished by its narrower (1.5–2 versus 2–5 mm) petals that are dark reddish purple to indigo (versus lavender or white). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The name Arabis (Boechera) divaricarpa has been applied to nearly every hybrid containing a genome derived from B. stricta. This presents a serious barrier to understanding the evolution of Boechera and also is contrary to the International Code of Botanical Nomenclature, because some names usually placed in synonymy (i.e., B. grahamii and B. brachycarpa) have priority at species level (M. D. Windham and I. A. Al-Shehbaz 2007b). To address this problem, we treat the following as distinct species: B. acutina, B. grahamii (= B. brachycarpa of R. D. Dorn 2001), and B. pratincola (all considered synonyms of A. divaricarpa by R. C. Rollins 1993), and B. calderi, B. elkoensis, and B. quebecensis (taxa described after 1993). Detailed comparison among these taxa are provided by Windham and Al-Shehbaz (2007, 2007b). The narrow concept of B. divaricarpa advocated here encompasses apomictic triploid populations containing three distinct genomes, one each derived from B. retrofracta, B. sparsiflora, and B. stricta. If the species is defined more broadly, the name B. grahamii has priority. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 366. | FNA vol. 7, p. 374. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Arabis atrorubens, Arabis atriflora, Arabis sparsiflora var. atrorubens | Arabis divaricarpa, Arabis stokesiae |
Name authority | (Suksdorf ex Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (A. Nelson) Á. Löve & D. Löve: Bot. Not. 128: 513. (1976) |
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