Boechera arcuata |
Boechera spatifolia |
|
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arching rockcress, elegant rockcress |
spoonleaf rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; short- to long-lived; sexual; caudex usually not woody (rarely with persistent, crowded leaf bases). |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
simple or few to several per caudex branch, arising from center of rosette near ground surface, 1.5–3.5(–5) dm, densely pubescent proximally, trichomes simple mixed with short-stalked ones, 2-rayed, 0.3–0.7 mm. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade narrowly oblanceolate, 1.5–3(–4) mm wide, margins entire, strongly ciliate at least along petiole, trichomes (simple), to 1 mm, surfaces glabrous or sparsely pubescent, trichomes simple and short-stalked, 2-rayed, 0.3–0.7 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
5–15(–20), often concealing stem proximally; blade auricles 0.5–1.5 mm, surfaces of distalmost leaves usually glabrous. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
10–30-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals usually white, rarely pale lavender, 3–3.7(–4) × 0.5–0.8 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
horizontal or slightly descending, curved or angled downward, 6–10(–15) mm, usually glabrous, rarely sparsely pubescent, trichomes spreading, simple. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
pendent, not appressed to rachis, rarely slightly secund, straight or gently curved, edges parallel, 3.3–5.7 cm × 1.2–1.8 mm; valves glabrous; ovules 90–126 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
biseriate, 0.7–0.9 × 0.5–0.6 mm; wing distal or, sometimes, absent, 0.05–0.1 mm wide. |
2n | = 14. |
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Boechera arcuata |
Boechera spatifolia |
|
Phenology | Flowering Mar–Jun. | Flowering May–Aug. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rocky slopes and gravelly soil in sagebrush, pinyon-juniper woodlands, open conifer forests and subalpine meadows |
Elevation | 300-1800 m (1000-5900 ft) | 1800-2800 m (5900-9200 ft) |
Distribution |
CA
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CO; NM
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Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera spatifolia is a sexual diploid that usually has been treated as a variety of Arabis (Boechera) fendleri but appears to be sufficiently distinct to warrant recognition at species level (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). There is little geographic overlap between the two, with B. spatifolia confined to the mountains of central Colorado and north-central New Mexico and B. fendleri ranging from western New Mexico and the Four Corners region through northern Arizona to southern Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 408. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis spatifolia, Arabis fendleri var. spatifolia, B. fendleri subsp. spatifolia, B. fendleri var. spatifolia |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Rydberg) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 84. (2006) |
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