Boechera arcuata |
Boechera repanda |
|
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arching rockcress, elegant rockcress |
Greene's rockcress, Yosemite rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; short- to long-lived; sexual; caudex usually not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, (1–)2–9 dm, densely pubescent proximally, trichomes short- to long-stalked, 2–6-rayed, 0.2–0.5 mm, rarely mixed with simple ones, to 1.5 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade broadly oblanceolate to obovate, 7–25(–50) mm wide, margins usually repand to coarsely dentate, rarely entire, not ciliate, surfaces moderately to densely pubescent, trichomes short- to long-stalked, 3–6-rayed, 0.2–0.5 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
(3–)8–30, not concealing stem; blade auricles absent, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
7–25-flowered, sometimes branched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
erect to ascending at anthesis; sepals glabrous or sparsely pubescent; petals white, 3.5–6 × 0.8–1 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
erect to divaricate-ascending, straight, 3–10 mm, glabrous or pubescent, trichomes spreading, usually 2–5-rayed, rarely simple. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
divaricate-ascending, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, (3.5–)5–13.5 cm × 2.5–4 mm; valves glabrous or pubescent throughout; ovules 34–50 per ovary; style 0.5–1.5 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 2.5–5(–6) × 2–3(–3.5) mm; wing continuous, 0.7–1.8 mm wide. |
2n | = 14. |
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Boechera arcuata |
Boechera repanda |
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Phenology | Flowering Mar–Jun. | Flowering Jun–Jul. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rock outcrops, talus, and gravelly soil in meadows and open pine forests |
Elevation | 300-1800 m (1000-5900 ft) | 1600-3500 m (5200-11500 ft) |
Distribution |
CA
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CA; NV
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Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera repanda is a sexual diploid that is distinguished from all other western North American species by its wide (10–50 mm), repand to coarsely dentate basal leaves. The form recognized as var. greenei appears to be a growth response to ecologically marginal habitats. Boechera repanda occurs sporadically in the Sierra Nevada and Transverse Ranges of California, and barely extends into the Carson Range in western Nevada. Arabis inamoena Greene (1911), not Greene (1908) is an illegitimate name, sometimes found in synonymy with Boechera repanda. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 402. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis repanda, Arabis repanda var. greenei |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (S. Watson) Al-Shehbaz: Novon 13: 388. (2003) |
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