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arching rockcress, elegant rockcress

Tulare County rock cress

Habit Perennials; usually long-lived; sexual; caudex often woody (well-developed). Perennials; usually long-lived; sexual; caudex woody, (often with persistent, crowded leaf bases).
Stems

usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally.

usually 2–5 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.2–0.8 dm, pubescent proximally, trichomes simple and short-stalked, 2- or 3-rayed, to 0.4 mm, glabrescent distally.

Basal leaves

blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm.

blade linear, 0.8–1.5 mm wide, margins entire, ciliate, trichomes (simple and spurred), to 0.8 mm, surfaces moderately pubescent, trichomes short-stalked, 2–4-rayed, 0.05–0.4 mm.

Cauline leaves

10–30(–45), often concealing stem proximally;

blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent.

2–4, not concealing stem;

blade auricles absent, surfaces of distalmost leaves pubescent.

Racemes

12–50(–70)-flowered, usually unbranched.

2–5-flowered, unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially);

pollen ellipsoid.

erect at anthesis;

sepals pubescent;

petals white, 3.5–5 × 0.7–1 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed.

erect to ascending, straight, 2–7 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched.

Fruits

usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm;

valves glabrous or trichomes relatively few, scattered;

ovules 90–250 per ovary;

style 0.01–0.5 mm.

erect to ascending, often appressed to rachis, not secund, straight, edges parallel, (1.3–)2–3.3 cm × 4–5 mm;

valves glabrous;

ovules 8–12 per ovary;

style 0.05–0.4 mm.

Seeds

uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm;

wing continuous, 0.1–0.2 mm wide.

uniseriate, 3–5 × 2.5–4.5 mm;

wing continuous, 0.8–2 mm wide distally.

Boechera arcuata

Boechera pygmaea

Phenology Flowering Mar–Jun. Flowering May–Jul.
Habitat Rocky hillsides and cliffs in pine forests and chaparral Barren flats of arkosic gravel
Elevation 300-1800 m (1000-5900 ft) 2400-3200 m (7900-10500 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Despite the distinctive morphology of Boechera pygmaea, there is evidence that it is capable of producing fertile hybrids with B. stricta. It is known only from Inyo and Tulare counties in the southern Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 365. FNA vol. 7, p. 401.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata Arabis pygmaea
Name authority (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) (Rollins) Al-Shehbaz: Novon 13: 388. (2003)
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