Boechera arcuata |
Boechera pusilla |
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arching rockcress, elegant rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; long-lived; (cespitose); apomictic; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 2–6 per caudex branch, arising from margin of rosette near ground surface, 0.5–2 dm, glabrous or sparsely pubescent proximally, trichomes simple and short-stalked, 2-rayed, to 0.2 mm, glabrous distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade linear-oblanceolate, 1–2.5 mm wide, margins entire, ciliate along petiole, trichomes (simple), 0.4–0.7 mm, surfaces usually sparsely pubescent, rarely glabrous, trichomes short-stalked, 2- or 3-rayed, 0.1–0.4 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
3–5, not concealing stem; blade auricles 0–0.2 mm, surfaces of distalmost leaves usually glabrous or, rarely, margins sparsely ciliate. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
6–13-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous or sparsely pubescent, trichomes spreading, 2-rayed; petals white to lavender, 4–5 × 1.5–1.8 mm, glabrous; pollen spheroid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
horizontal to divaricate-descending, straight or slightly curved downward, 2–5 mm, glabrous. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
horizontal or divaricate-descending, not appressed to rachis, secund, straight, edges parallel, 1.6–3.2 cm × 1.5–2 mm; valves glabrous; ovules 20–32 per ovary; style 0.1–0.4 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 1.2–1.5 × 0.8–0.9 mm; not winged or with distal wing 0.05–0.1 mm wide. |
Boechera arcuata |
Boechera pusilla |
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Phenology | Flowering Mar–Jun. | Flowering May–Jun. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Cracks and crevices of granitic rock outcrops |
Elevation | 300-1800 m (1000-5900 ft) | 2400-2500 m (7900-8200 ft) |
Distribution |
CA
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WY |
Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Morphological evidence suggests that Boechera pusilla is an apomictic species that arose through hybridization between B. lemmonii and B. pendulina; it is known only from the type locality in southwestern Wyoming. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 400. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis pusilla |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Rollins) Dorn: Vasc. Pl. Wyoming ed. 3, 376. (2001) |
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