Boechera arcuata |
Boechera pendulina |
|
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arching rockcress, elegant rockcress |
nodding rock cress, rabbit-ear rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; short- to long-lived; sexual or apomictic; caudex sometimes woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 2–6 per caudex branch, arising from margin of rosette near ground surface, or arising laterally proximal to sterile shoots, 0.6–3(–3.7) dm, sparsely to densely pubescent proximally, trichomes simple, 0.3–0.8 mm, glabrous distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade oblanceolate to obovate, 1.5–6 mm wide, margins usually entire, rarely dentate, ciliate throughout, trichomes (usually simple), 0.4–1 mm, surfaces pubescent, trichomes simple and short- and long-stalked, 2-rayed, 0.3–0.8 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
2–10(–13), not concealing stem; blade auricles absent or, rarely, to 0.7 mm, surfaces of distalmost leaves glabrous or margins sparsely ciliate. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
4–14-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-ascending at anthesis; sepals usually sparsely pubescent, rarely glabrous; petals whitish to pale lavender, 4–6 × 1–1.5 mm, glabrous; pollen ellipsoid or spheroid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
divaricate-ascending to horizontal, curved or angled downward, 3–7(–10) mm, usually glabrous, rarely with some simple trichomes. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
widely pendent, not appressed to rachis, not secund, curved to nearly straight, edges parallel, 2.2–4 cm × 1.2–2.1 mm; valves glabrous; ovules 40–70(–90) per ovary; style 0.1–0.3(–0.5) mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
biseriate, 0.9–1.2 × 0.6–0.9 mm, usually not winged. |
2n | = 14, 21. |
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Boechera arcuata |
Boechera pendulina |
|
Phenology | Flowering Mar–Jun. | Flowering Apr–Jun. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rock outcrops, open gravelly flats and hillsides in sagebrush, pinyon-juniper, mountain mahogany, open conifer forests |
Elevation | 300-1800 m (1000-5900 ft) | 1600-3100 m (5200-10200 ft) |
Distribution |
CA
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AZ; CA; CO; NV; UT; WY
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Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although included in Arabis (Boechera) demissa by some authors (e.g., S. L. Welsh et al. 2003; N. H. Holmgren 2005b), B. pendulina is readily distinguished from that species (see M. D. Windham and I. A. Al-Shehbaz 2006 for detailed comparison). Typical collections are sexual diploids, whereas the type of var. russeola is a triploid apomict; further study is needed to determine if the two are conspecific. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 395. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis pendulina, Arabis demissa var. russeola, Arabis diehlii, Arabis pendulina var. russeola, Arabis setulosa, B. demissa var. pendulina, B. demissa var. russeola, B. pendulina var. russeola |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Greene) W. A. Weber: Phytologia 51: 370. (1982) |
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