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arching rockcress, elegant rockcress

Koehler's rockcress, shrubby rock-cress

Habit Perennials; usually long-lived; sexual; caudex often woody (well-developed). Perennials; long-lived; sexual; caudex woody (with persistent, peg-like leaf bases).
Stems

usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally.

usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 0.8–4.5 dm, pubescent proximally, trichomes stalked, 2–4-rayed, to 0.5 mm, mixed with simple ones, to 1 mm, glabrous distally.

Basal leaves

blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm.

blade narrowly oblanceolate, 1–3 mm wide, margins entire, ciliate proximally, trichomes (simple), to 1 mm, surfaces moderately to sparsely pubescent, trichomes stalked, 2–4-rayed, 0.1–0.3(–0.5) mm.

Cauline leaves

10–30(–45), often concealing stem proximally;

blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent.

3–17(–30), rarely concealing stem;

blade auricles 0.5–2.5 mm, surfaces of distalmost leaves glabrous.

Racemes

12–50(–70)-flowered, usually unbranched.

6–20(–35)-flowered, usually unbranched.

Flowers

ascending at anthesis;

sepals pubescent;

petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially);

pollen ellipsoid.

ascending at anthesis;

sepals pubescent;

petals deep purple to magenta, 8–12 × 2.5–4 mm, glabrous;

pollen ellipsoid.

Fruiting pedicels

usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed.

divaricate-ascending to horizontal, straight, 10–18 mm, glabrous or sparsely pubescent, trichomes spreading, simple and 2-rayed.

Fruits

usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm;

valves glabrous or trichomes relatively few, scattered;

ovules 90–250 per ovary;

style 0.01–0.5 mm.

divaricate-ascending to widely pendent, not appressed to rachis, not secund, usually curved, rarely straight, edges parallel, 5–7.5 cm × 1.8–2.5 mm;

valves glabrous;

ovules 58–94 per ovary;

style 0.05–0.2 mm.

Seeds

uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm;

wing continuous, 0.1–0.2 mm wide.

uniseriate, 1.3–1.8 × 1.2–1.5 mm;

wing continuous, 0.1–0.2 mm wide.

Boechera arcuata

Boechera koehleri

Phenology Flowering Mar–Jun. Flowering Apr–May.
Habitat Rocky hillsides and cliffs in pine forests and chaparral Serpentine and limestone outcrops
Elevation 300-1800 m (1000-5900 ft) 100-500 m (300-1600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Boechera koehleri var. koehlerei is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 365. FNA vol. 7, p. 386.
Parent taxa Brassicaceae > tribe Boechereae > Boechera Brassicaceae > tribe Boechereae > Boechera
Sibling taxa
B. acutina, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. koehleri, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
B. acutina, B. arcuata, B. atrorubens, B. bodiensis, B. breweri, B. burkii, B. calderi, B. californica, B. canadensis, B. cascadensis, B. cobrensis, B. collinsii, B. consanguinea, B. constancei, B. covillei, B. crandallii, B. cusickii, B. davidsonii, B. dentata, B. depauperata, B. dispar, B. divaricarpa, B. drepanoloba, B. duchesnensis, B. elkoensis, B. evadens, B. falcatoria, B. falcifructa, B. fecunda, B. fendleri, B. fernaldiana, B. formosa, B. fructicosa, B. glareosa, B. glaucovalvula, B. goodrichii, B. gracilenta, B. gracilipes, B. grahamii, B. gunnisoniana, B. harrisonii, B. hastatula, B. hoffmannii, B. holboellii, B. horizontalis, B. howellii, B. inyoensis, B. johnstonii, B. laevigata, B. languida, B. lasiocarpa, B. lemmonii, B. lignifera, B. lincolnensis, B. lyallii, B. macounii, B. microphylla, B. missouriensis, B. nevadensis, B. ophira, B. oxylobula, B. paddoensis, B. pallidifolia, B. parishii, B. pauciflora, B. paupercula, B. peirsonii, B. pendulina, B. pendulocarpa, B. perennans, B. perstellata, B. pinetorum, B. pinzliae, B. platysperma, B. polyantha, B. porphyrea, B. pratincola, B. puberula, B. pulchra, B. pusilla, B. pygmaea, B. quebecensis, B. rectissima, B. repanda, B. retrofracta, B. rigidissima, B. rollei, B. rollinsiorum, B. rubicundula, B. saximontana, B. schistacea, B. serotina, B. serpenticola, B. shevockii, B. shockleyi, B. sparsiflora, B. spatifolia, B. stricta, B. subpinnatifida, B. suffrutescens, B. texana, B. tiehmii, B. tularensis, B. ultra-alsa, B. villosa, B. williamsii, B. xylopoda, B. yorkii
Synonyms Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata Arabis koehleri, Arabis arbuscula, Arabis koehleri var. stipitata
Name authority (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) (Howell) Al-Shehbaz: Novon 13: 386. (2003)
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