Boechera arcuata |
Boechera gunnisoniana |
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arching rockcress, elegant rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; usually long-lived; (cespitose); sexual; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 2–6 per caudex branch, arising from margin of rosette near ground surface, 0.8–2(–2.5) dm, densely pubescent proximally, trichomes short-stalked, 2–4-rayed, 0.08–0.3 mm, mixed with fewer simple ones, usually sparsely pubescent distally, rarely glabrous. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade linear-oblanceolate, 1–3(–4) mm wide, margins entire, ciliate proximally, trichomes to 0.8 mm, surfaces densely pubescent, trichomes short-stalked, 3–6-rayed, 0.08–0.3 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
3–9, not concealing stem; blade auricles 0.2–1 mm, surfaces of distalmost leaves glabrous or sparsely pubescent. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
7–15-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals white or lavender, 4–6 × 1–2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
divaricate, straight or slightly curved, 4–7 mm, glabrous or sparsely pubescent, trichomes appressed, branched. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
horizontal to divaricate-descending, not appressed to rachis, not secund, straight to slightly curved, edges parallel, 2.5–4 cm × 1–1.5 mm; valves glabrous; ovules 36–54 per ovary; style 0.1–0.2 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 1–1.2 × 0.8–1 mm; wing continuous, 0.1–0.15 mm wide. |
2n | = 14. |
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Boechera arcuata |
Boechera gunnisoniana |
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Phenology | Flowering Mar–Jun. | Flowering May–Jun. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rocky slopes and knolls with sagebrush |
Elevation | 300-1800 m (1000-5900 ft) | 2100-2700 m (6900-8900 ft) |
Distribution |
CA
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CO |
Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera gunnisoniana is most similar to B. glareosa; it is distinguished from that species by having trichomes smaller (0.08–0.3 versus 0.3–0.6 mm) and more highly divided (mostly 2–4-rayed versus simple and 2-rayed) proximally on stems, and fewer (36–54 versus 50–80) ovules per ovary. The two species are allopatric, with B. glareosa known only from eastern Utah and extreme northwestern Colorado and B. gunnisoniana apparently endemic to Gunnison County in west-central Colorado. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 382. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis gunnisoniana |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Rollins) W. A. Weber: Phytologia 51: 370. (1982) |
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