Boechera arcuata |
Boechera formosa |
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arching rockcress, elegant rockcress |
beautiful rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 1 per caudex branch, arising from center of leaf tuft, above ground surface on woody base, 2–5.5 dm, densely pubescent proximally, trichomes short-stalked, 4–7-rayed, 0.1–0.4 mm, similarly pubescent distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade linear to linear-oblanceolate, 2–3(–4) mm wide, margins entire, sometimes ciliate near petiole base, trichomes (simple or 2-rayed), to 0.8 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.3 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
7–18, concealing stem proximally; blade auricles absent, surfaces of distalmost leaves densely pubescent. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
6–26-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate at anthesis; sepals pubescent; petals white to pale lavender, 8–18 × 2.5–4.4(–5.5) mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
horizontal to descending, usually recurved, 5–10(–20) mm, pubescent, trichomes appressed, branched. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
divaricate-descending to reflexed, usually not appressed to rachis, rarely somewhat secund, straight, edges parallel, 4.3–7 cm × 1.6–3 mm; valves pubescent throughout; ovules 72–98 per ovary; style ca. 0.1 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
biseriate, 1.2–1.6 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
2n | = 14. |
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Boechera arcuata |
Boechera formosa |
|
Phenology | Flowering Mar–Jun. | Flowering Apr–Jun. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rocky slopes and sandy soil in blackbrush, sagebrush, and pinyon-juniper communities |
Elevation | 300-1800 m (1000-5900 ft) | 1300-1900 m (4300-6200 ft) |
Distribution |
CA
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AZ; CO; NM; UT; WY
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Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although Boechera formosa is usually subsumed within Arabis (Boechera) pulchra, the two species are easily separated. Boechera formosa has white to lavender (versus mostly purple) petals, fruiting pedicels that are not abruptly recurved at the base, narrower (1.6–3 versus 2.5–4 mm wide), mostly non-appressed fruits, and narrower (1–1.2 versus 1.5–2.2 mm wide) seeds. The two taxa are separated by more than 500 km, with B. formosa found only on the Colorado Plateau and B. pulchra, in the strict sense, restricted to southern California, western Nevada, and northwestern Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 379. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis formosa, Arabis pulchra var. pallens, B. pulchra subsp. pallens, B. pulchra var. pallens |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 68. (2006) |
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