Boechera arcuata |
Boechera duchesnensis |
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arching rockcress, elegant rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; usually long-lived; apomictic; caudex often woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from near ground surface, 1.5–4.5 dm, densely pubescent proximally, trichomes short-stalked, 2–7-rayed, 0.2–0.5 mm (sometimes mixed with some larger trichomes proximally), sparsely pubescent distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade linear-oblanceolate, 2–5 mm wide, margins entire or denticulate, ciliate at petiole base, trichomes (simple), to 1 mm, surfaces densely pubescent, trichomes short-stalked, 4–8-rayed, 0.15–0.4 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
3–8, not concealing stem; blade auricles 0.7–2 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
11–22-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-ascending at anthesis; sepals pubescent; petals whitish to pale lavender, 7–10 × 1.5–3 mm, glabrous; pollen spheroid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
horizontal, straight or slightly curved, 7–14 mm, pubescent, trichomes appressed, branched. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
horizontal, not appressed to rachis, not secund, usually straight, edges parallel, 3.5–5 cm × 1.7–2 mm; valves glabrous proximally, pubescent distally; ovules 72–92 per ovary; style 0.1–0.3 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
sub-biseriate, 1.1–1.3 × 0.7–1 mm; wing continuous, ca. 0.1 mm wide. |
Boechera arcuata |
Boechera duchesnensis |
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Phenology | Flowering Mar–Jun. | Flowering May. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Sandy soil on rocky slopes, mostly in pinyon-juniper woodlands |
Elevation | 300-1800 m (1000-5900 ft) | 1300-1800 m (4300-5900 ft) |
Distribution |
CA
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CO; NM; UT |
Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological and isozyme analyses indicate that Boechera duchesnensis is an apomictic species that arose through hybridization between B. formosa and B. pallidifolia (M. D. Windham and Allphin, unpubl.). It is easily distinguished from those species by having fruits that are glabrous proximally and sparsely pubescent distally. It shares this distinctive fruit pubescence with B. xylopoda, an apomictic hybrid between B. perennans and B. pulchra (see Windham and I. A. Al-Shehbaz 2007b for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 375. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis pulchra var. duchesnensis, B. pulchra var. duchesnensis |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Rollins) Windham: Harvard Pap. Bot. 12: 239. (2007) |
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