Boechera arcuata |
Boechera constancei |
|
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arching rockcress, elegant rockcress |
Constance's Rock cress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; long-lived; sexual; caudex woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 1 per caudex branch, arising from center of rosette, elevated above ground surface on woody base, 1.2–3 dm, glabrous throughout. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade narrowly oblanceolate, 1.5–4 mm wide, margins entire, ciliate, trichomes (simple, mixed with fewer stalked, 2-rayed), 0.3–0.8 mm, surfaces glabrous. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
6–12, not concealing stem; blade auricles absent, surfaces of distalmost leaves glabrous. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
5–15-flowered, unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-ascending at anthesis; sepals glabrous; petals creamy white, 6–8 × 1.5–2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
arched, strongly recurved proximally, 4–12 mm, glabrous. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
pendent, not appressed to rachis, usually secund, straight or slightly curved, edges often somewhat undulate (not parallel), 4–7.5 cm × 3–3.5 mm; valves glabrous; ovules 18–28 per ovary; style 2.5–5 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 3–4 × 2.5–3 mm; wing continuous, 0.5–1 mm wide. |
2n | = 14. |
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Boechera arcuata |
Boechera constancei |
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Phenology | Flowering Mar–Jun. | Flowering May. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Steep ridges, serpentine slopes |
Elevation | 300-1800 m (1000-5900 ft) | 1200-1900 m (3900-6200 ft) |
Distribution |
CA
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CA |
Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera constancei was originally treated as a variety of Arabis (Boechera) suffrutescens; it is easily separated from that species by its non-auriculate cauline leaf blades, longer (6–8 versus 4.5–6 mm) petals that are creamy white (versus purple or rose-tipped), much longer style (2.5–5 versus 0.4–1.2 mm), and exerted anthers. This distinctive sexual diploid is thus far known only from the west slope of the Sierra Nevada in Plumas County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 371. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis constancei, Arabis suffrutescens var. perstylosa |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Rollins) Al-Shehbaz: Novon 13: 384. (2003) |
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