Boechera arcuata |
Boechera consanguinea |
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arching rockcress, elegant rockcress |
Four Corners rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Biennials or perennials; short-lived; apomictic; caudex present or absent. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 1.5–5 dm, densely pubescent proximally, trichomes short-stalked, 2–6-rayed, 0.2–0.5 mm, glabrous or sparsely pubescent distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade oblanceolate, 2–10 mm wide, margins usually dentate, sometimes ciliate near petiole base, trichomes (usually spurred), to 0.7 mm, surfaces moderately to densely pubescent, trichomes short-stalked, 4–8-rayed, 0.1–0.4 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
15–36, sometimes concealing stem proximally; blade auricles 1–3 mm, surfaces of distalmost leaves sparsely pubescent. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
20–55-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
divaricate-descending at anthesis; sepals pubescent; petals pale lavender, 5–8.5 × 1–2 mm, glabrous; pollen spheroid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
reflexed to divaricate-descending, usually curved downward, 8–14 mm, usually pubescent, rarely glabrous, trichomes subappressed, branched. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
reflexed to pendent, rarely appressed to rachis, not secund, straight or slightly curved, edges parallel, 4–6 cm × 1–2 mm; valves glabrous; ovules 100–128 per ovary; style 0.05–0.5 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
biseriate to sub-biseriate, mature seeds not seen. |
Boechera arcuata |
Boechera consanguinea |
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Phenology | Flowering Mar–Jun. | Flowering May. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rocky slopes and sandy soil in ponderosa pine, pinyon-juniper, and sagebrush communities |
Elevation | 300-1800 m (1000-5900 ft) | 1900-2500 m (6200-8200 ft) |
Distribution |
CA
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CO; NM; UT
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Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Morphological evidence suggests that Boechera consanguinea is an apomictic species that arose through hybridization between B. fendleri and B. retrofracta. It is most similar to B. goodrichii, another apomictic hybrid involving B. retrofracta (see M. D. Windham and I. A. Al-Shehbaz 2007 for detailed comparison). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 371. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis consanguinea, Arabis holboellii var. consanguinea |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 261. (2007) |
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