Boechera arcuata |
Boechera atrorubens |
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arching rockcress, elegant rockcress |
black-flower rockcress, dark-red-flower rockcress, purple sickle-pod rock-cress, sicklepod rockcress |
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Habit | Perennials; usually long-lived; sexual; caudex often woody (well-developed). | Perennials; usually short-lived; sexual; caudex not woody. |
Stems | usually 1 per caudex branch, arising from center of rosette, elevated on woody base or from ground surface, (2–)3–8 dm, densely pubescent proximally, trichomes short-stalked, 2-rayed and simple, to 1 mm, pubescent distally. |
usually 1 per caudex branch, arising from center of rosette near ground surface, 0.8–6 dm, sparsely pubescent proximally, trichomes short-stalked and simple, (scattered), 2-rayed, 0.1–0.15 mm, glabrous distally. |
Basal leaves | blade linear to oblanceolate, 2–7(–12) mm wide, margins usually entire, rarely denticulate, ciliate along petiole, trichomes to 1.5 mm, surfaces densely pubescent, trichomes usually short-stalked, 2–5-rayed (rarely some simple), 0.4–0.8 mm. |
blade oblanceolate, 4–10 mm wide, margins usually dentate, not ciliate, surfaces densely pubescent, trichomes short-stalked, 3- or 4-rayed, 0.1–0.2 mm. |
Cauline leaves | 10–30(–45), often concealing stem proximally; blade auricles 2–5(–6) mm, surfaces of distalmost leaves pubescent. |
7–25, concealing stem proximally; blade auricles 1–4 mm, surfaces of distalmost leaves glabrous. |
Racemes | 12–50(–70)-flowered, usually unbranched. |
6–20-flowered, usually unbranched. |
Flowers | ascending at anthesis; sepals pubescent; petals purple, 9–14 × 2–4 mm, glabrous or sparsely pubescent (trichomes abaxially); pollen ellipsoid. |
ascending at anthesis; sepals pubescent; petals dark reddish purple (drying indigo), 7–9 × 1.5–2 mm, glabrous; pollen ellipsoid. |
Fruiting pedicels | usually divaricate-ascending, rarely horizontal, gently recurved or straight, 8–22 mm, pubescent, trichomes subappressed, 2–4-rayed. |
ascending, usually straight, 5–10 mm, glabrous or pubescent, trichomes spreading, simple. |
Fruits | usually divaricate-ascending, rarely horizontal, not appressed to rachis, not secund, usually curved, edges parallel, (6–)8–13 cm × 1.5–2.2 mm; valves glabrous or trichomes relatively few, scattered; ovules 90–250 per ovary; style 0.01–0.5 mm. |
ascending, not appressed to rachis, not secund, curved or straight, edges parallel, (4–)6–12 cm × 1.7–2.2 mm; valves glabrous; ovules 80–100 per ovary; style 0.2–0.5 mm. |
Seeds | uniseriate or sub-biseriate, 1.5–1.7 × 1–1.2 mm; wing continuous, 0.1–0.2 mm wide. |
uniseriate, 1.2–1.7 × 0.9–1.2 mm; wing lateral and distal, 0.1–0.2 mm wide. |
Boechera arcuata |
Boechera atrorubens |
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Phenology | Flowering Mar–Jun. | Flowering Apr–May. |
Habitat | Rocky hillsides and cliffs in pine forests and chaparral | Rocky summits and sandy loam on sagebrush slopes |
Elevation | 300-1800 m (1000-5900 ft) | ca. 600 m (ca. 2000 ft) |
Distribution |
CA
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OR; WA
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Discussion | Although usually treated as a variety of Arabis (Boechera) sparsiflora (e.g., R. C. Rollins 1993), B. arcuata is easily distinguished from that species by having rachises and fruiting pedicels pubescent with subappressed, 2–4-rayed trichomes and a geographic range limited to southern and western California. By contrast, B. sparsiflora has rachises and fruiting pedicels with spreading, usually simple trichomes (sometimes glabrescent) and an allopatric distribution north and east of the Sierra Nevada. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Boechera atrorubens is often treated as a variety of B. sparsiflora (e.g., R. C. Rollins 1993), it is readily separated from that species by having proximal stems sparsely (versus densely) pubescent with much smaller (0.15 versus 1.5 mm) trichomes. The two taxa rarely grow in proximity and, in areas where they are sympatric, B. atrorubens is further distinguished by its narrower (1.5–2 versus 2–5 mm) petals that are dark reddish purple to indigo (versus lavender or white). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 365. | FNA vol. 7, p. 366. |
Parent taxa | Brassicaceae > tribe Boechereae > Boechera | Brassicaceae > tribe Boechereae > Boechera |
Sibling taxa | ||
Synonyms | Streptanthus arcuatus, Arabis holboellii var. arcuata, Arabis maxima, Arabis sparsiflora var. arcuata | Arabis atrorubens, Arabis atriflora, Arabis sparsiflora var. atrorubens |
Name authority | (Nuttall) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) | (Suksdorf ex Greene) Windham & Al-Shehbaz: Harvard Pap. Bot. 11: 64. (2006) |
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