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hammock fern

chain fern family, deer fern family

Habit Plants perennial, mostly terrestrial, occasionally on rock or epiphytic.
Stems

creeping to suberect or ascending, sometimes climbing [rarely arborescent], slender to stout, dictyostelic, scaly.

Leaves

monomorphic or dimorphic, large and coarse, generally greater than 30 cm, often exceeding 1 m. Petiole not articulate, generally more than 2 vascular bundles arranged in arc, generally scaly at least at base.

Blade

often anthocyanic (reddish) when young, pinnatifid [rarely simple] to pinnate-pinnatifid or 2-pinnate [rarely decompound], glabrous or occasionally bearing scales or capitate glands.

Veins

of sterile leaves generally free, rarely anastomosing, veins of fertile leaves united to form sorus-bearing secondary vein parallel to costa or costule (vascular commisure), sometimes anastomosing further.

Sori

elongate along secondary vein;

indusia present [rarely absent], opening along costal side of fertile vein, frequently hidden by dehisced sporangia;

sporangial stalk of 3 rows of cells.

Spores

monolete, reniform;

perine present, variously ornamented.

Gametophytes

green, cordate, sometimes bearing capitate hairs, antheridia and archegonia borne on lower surface.

Rachis

frequently grooved adaxially.

Blechnum occidentale

Blechnaceae

Distribution
from FNA
FL; GA; LA; TX; Mexico; Central America; South America; West Indies
Mostly tropical and south temperate (except Woodwardia, which is north temperate)
[BONAP county map]
Discussion

Varieties 2 (1 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Circumscription of genera is controversial, especially as to placement of those species now included in Blechnum. Characteristics holding the family together include the anastomoses of veins along the axes of the blade to form a series of areoles or a single continuous vein along which the sorus is borne, elongate sori with indusia opening toward midvein, bilateral spores, and chromosome base numbers of generally x = 28–36. Relationships of the family with both dryopteroid and athyrioid ferns have been suggested.

Stenochlaena tenuifolia (Desvaux) T. Moore, native to the Old World, was reported as escaped from cultivation in the 1930s in southern Florida; it has not been collected there recently. It is distinguished by having climbing stems and by having contracted, 2-pinnate fertile leaves with sporangia covering the abaxial surface.

Genera ca. 10, species ca. 250 (2 genera, 6 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Sori continuous along length of costa or costule; veins of sterile leaves free.
Blechnum
1. Sori distinct, in chainlike rows along costa or costule; veins of sterile leaves anastomosing at least along costae and costules.
Woodwardia
Source FNA vol. 2. FNA vol. 2, p. 223. Author: Raymond B. Cranfill.
Parent taxa Blechnaceae > Blechnum
Sibling taxa
B. serrulatum, B. spicant
Subordinate taxa
B. occidentale var. minor
Blechnum, Woodwardia
Name authority Linnaeus: Sp. Pl., ed. 2 2: 1524. (1763) C. Presl
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