Bistorta vivipara |
Bistorta |
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alpine bistort, serpent-grass, viviparous bistort |
bistort |
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Habit | Plants (2–)8–30(–45) cm; rhizomes sometimes contorted. | Herbs, perennial; roots fibrous, rhizomatous. | ||||||||||||
Stems | 1–2(–3). |
erect, simple, glabrous. |
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Leaves | ocrea brown, cylindric, 4–22(–27) mm, margins strongly oblique, glabrous; petiole attached to sheath 6–20(–45) mm, unwinged distally, 5–110(–200) mm; blade linear to lanceolate or oblong-ovate, 1–8(–10) × 0.5–1.7(–2.3) cm, base cuneate to rounded or cordate, often asymmetric, margins entire, usually revolute, not wavy, apex obtuse to acute, abaxial face pubescent with whitish or brownish hairs, glaucous, adaxial face glabrous, not glaucous; cauline leaves 2–4, petiolate proximally, sessile distally, gradually reduced distally, blade linear-lanceolate to linear. |
mostly basal, some cauline, alternate, petiolate or sessile; ocrea persistent or disintegrating with age and deciduous entirely or distally, chartaceous; blade linear or lanceolate to elliptic, oblong-ovate, or ovate, margins entire or obscurely and irregularly repand. |
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Inflorescences | 1, narrowly elongate-cylindric, (15–)20–90 × 4–10 mm, usually bearing pink to brown or purple pyriform bulblets proximally and sterile flowers distally; peduncle 1–5 cm. |
terminal, spikelike. |
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Pedicels | ascending or spreading, (1–)2–5 mm. |
present. |
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Flowers | 1–2 per ocreate fascicle; perianth greenish proximally, usually white or pink distally, rarely red; tepals obovate, 2.1–4 mm, apex obtuse to acute; stamens included or exserted, some or all often poorly developed; anthers reddish to purple. |
bisexual, 1–2 per ocreate fascicle, base not stipelike; perianth nonaccrescent, white, greenish white, pink, or purplish pink, rarely red, campanulate, glabrous; tepals 5, connate proximally ca. 1/5 their length, petaloid, monomorphic or slightly dimorphic, outer larger than inner; stamens 5–8, sometimes poorly developed; filaments distinct or connate basally, outer ones sometimes adnate to perianth tube, glabrous; anthers yellow, pink, red, purple, or blackish, ovate to elliptic; styles 3, erect or spreading, distinct or connate proximally; stigmas 2–3, capitate. |
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Achenes | rarely produced, dark brown, 2.2–3.3 × 0.9–1.5 mm, dull, granular. |
included or exserted, brown to dark brown, unwinged, 3-gonous, glabrous. |
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Seeds | embryo curved. |
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x | = 11, 12. |
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2n | = 96, 120. |
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Bistorta vivipara |
Bistorta |
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Phenology | Flowering Jun–Sep. | |||||||||||||
Habitat | Moist to wet spruce or mixed woods along shorelines, moist subalpine woods and meadows, alpine meadows, heaths, nutrient-rich sites | |||||||||||||
Elevation | 0-4000 m (0-13100 ft) | |||||||||||||
Distribution |
AK; AZ; CO; ID; ME; MI; MN; MT; NH; NM; NV; OR; SD; UT; VT; WA; WY; AB; BC; LB; MB; NB; NT; NU; ON; QC; SK; YT; SPM; Greenland; Europe; Asia
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Arctic and temperate North America; Europe; Asia |
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Discussion | Bistorta vivipara is highly variable morphologically and cytologically. Robust plants with large leaves, compact spikes, and persistent bulblets have been named subsp. macounii. Abortion of stamens, production of bulblets, and the rarity of fruits suggest that reproduction is largely asexual; fruits and seedlings are produced rarely (N. Söyrinki 1989). B. Jonsell and T. Karlsson (2000+, vol. 1) summarized chromosome numbers that include 2n = 66, ca. 77, ca. 80, 88, 99, ca. 100, 110, 120, and ca. 132. A. E. Porsild and W. J. Cody (1980) reported that indigenous peoples of the circumpolar region eat the starchy, slightly astringent rootstocks raw or cooked, and preserve them in seal oil or by freezing. E. Hultén (1968) reported that the rootstocks taste like almonds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 50 (4 in the flora). Bistorta often is included in Polygonum in the broad sense or in Persicaria. It is accepted here as a distinct genus based on habit, morphology, and anatomy (K. Haraldson 1978; L.-P. Ronse Decraene and J. R. Akeroyd 1988). In the species of the flora area, the base of the petiole forms a long, tubular sheath distal to the node from which the leaf arises and proximal to the point of divergence of the petiole. Distal to the sheath is the ocrea, which usually is darker and thinner. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 595. | FNA vol. 5, p. 594. | ||||||||||||
Parent taxa | Polygonaceae > subfam. Polygonoideae > Bistorta | Polygonaceae > subfam. Polygonoideae | ||||||||||||
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Subordinate taxa | ||||||||||||||
Synonyms | Polygonum viviparum, B. vivipara subsp. macounii, Persicaria vivipara, Polygonum viviparum var. macounii | Polygonum unranked B. | ||||||||||||
Name authority | (Linnaeus) Delarbre: Fl. Auvergne ed. 2, 2: 516. (1800) | (Linnaeus) Scopoli: Meth. Pl., 24. (1754) | ||||||||||||
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