Bidens tripartita |
Bidens trichosperma |
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bident à trois divisions, three-lobed beggar-ticks, three-lobed beggars-ticks, three-part beggarticks, threelobe beggarticks, tickseed |
crowned beggar-ticks, tall swamp marigold |
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Habit | Annuals, (5–)20–70(–200) cm. | Annuals (biennials), (15–)30–50(–150+) cm. |
Leaves | sessile or petioles 5–15(–35+) mm (± winged); blades ± elliptic to ovate or lanceolate, 40–80(–150+) × 15–40(–60+) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices acute to acuminate, faces glabrous or hirtellous. |
petioles 5–20+ mm; blades ± deltate to ovate overall, 40–75(–150) × 10–45(–130) mm, usually laciniately 1–2-pinnatisect, ultimate lobes (3–)5–7+, oblanceolate or lanceolate to linear, (5–)10–30(–80+) × (1–)4–8(–12+) mm, bases cuneate, ultimate margins incised, dentate, serrate, or entire, little, if at all, ciliate, apices acute to attenuate, faces glabrous or ± hirtellous to strigillose. |
Peduncles | 10–40(–80) mm. |
20–150 mm. |
Involucres | campanulate to hemispheric or broader, (4–)5–7(–12) × (3–)6–12(–15+) mm. |
hemispheric or broader, 4–6(–8) × 6–12 mm. |
Ray florets | usually 0, sometimes 1–5; laminae orange yellowish, 4–8 mm. |
(7–)8–9; laminae golden yellow, 10–30 mm. |
Disc florets | (5–)20–60(–150+); corollas pale yellow to orange, (2–)3–4 mm (gradually ampliate, anthers usually pale). |
40–60(–80+); corollas yellow, 3–5 mm. |
Phyllaries | (6–)7–8(–13), elliptic-ovate to lance-ovate, (4–)6–9(–12) mm. |
6–8(–10), oblong, 3–8 mm. |
Calyculi | of (2–)6–7(–10) ± spreading, oblanceolate or lanceolate to linear, ± foliaceous bractlets or bracts 7–35(–60) mm, margins (entire or serrate) sometimes sparsely ciliate, abaxial faces hispidulous near bases, distally glabrous. |
of (6–)8(–11), ascending to spreading, spatulate to linear, sometimes ± foliaceous bractlets or bracts 3–10(–18) mm, margins sometimes ciliate, abaxial faces glabrous. |
Heads | borne singly or in 2s or 3s. |
usually in open, ± corymbiform arrays. |
Cypselae | blackish to purplish or brown, ± flattened, sometimes weakly 3(–4)-angled, usually cuneate to linear, outer (3–)6–7(–10) mm, inner (4–)6–9(–11) mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces ± 1-nerved, usually smooth, seldom notably tuberculate, glabrous or sparsely strigillose; pappi 0, or of (1–)3–3(–4+) ± erect to spreading, retrorsely barbed awns (0.2–)2–3(–6) mm. |
blackish or brown, flattened, narrowly cuneate, outer 3–6 mm, inner 5–9 mm (lengths mostly 2.5–4 times widths), margins antrorsely barbed or ciliate, apices ± truncate, faces obscurely 1-nerved, sometimes tuberculate, glabrous or sparsely hispidulous; pappi of 2 erect, ± patently barbed awns or scales (0.4–)1–2.5(–4) mm. |
2n | = 48. |
= 24. |
Bidens tripartita |
Bidens trichosperma |
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Phenology | Flowering Aug–Oct. | Flowering Aug–Oct. |
Habitat | Marshes and other wet sites | Marshes, estuaries |
Elevation | 10–1700 m (0–5600 ft) | 0–300 m (0–1000 ft) |
Distribution |
AK; AL; CA; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
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AL; AR; CT; DE; FL; GA; IA; IL; IN; KY; MA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; PA; RI; SC; SD; TN; VA; WI; WV; ON; QC
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Discussion | Plants with cypsela mid-nerves strongly developed (cypselae more or less strongly 4-angled and, often, tuberculate) that are treated below as Bidens connata have been included in B. tripartita, perhaps rightly so. And some botanists have included (or advocated inclusion of) B. eatonii, B. heterodoxa, and/or B. infirma in B. tripartita, as well, perhaps rightly so. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants here called Bidens trichosperma have long been known as B. coronata (Linnaeus) Britton (or Britton ex Sherff). Alas, Britton’s B. coronata (1913) is a later homonym of B. coronata Fischer ex Colla (1834) and cannot be used. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 217. | FNA vol. 21, p. 213. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens |
Sibling taxa | ||
Synonyms | B. comosa | Coreopsis trichosperma, B. coronata var. brachyodonta, B. coronata var. tenuiloba, B. coronata var. trichosperma |
Name authority | Linnaeus: Sp. Pl. 2: 831. (1753) | (Michaux) Britton: Bull. Torrey Bot. Club 20: 281. (1893) |
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