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bident à trois divisions, three-lobed beggar-ticks, three-lobed beggars-ticks, three-part beggarticks, threelobe beggarticks, tickseed

bident poilu, cobbler's pegs, common beggar-ticks, hairy beggar-ticks, Spanish needle

Habit Annuals, (5–)20–70(–200) cm. Annuals [perennials], (10–)30–60(–180+)[–250] cm.
Leaves

sessile or petioles 5–15(–35+) mm (± winged);

blades ± elliptic to ovate or lanceolate, 40–80(–150+) × 15–40(–60+) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices acute to acuminate, faces glabrous or hirtellous.

petioles 10–30(–70) mm;

blades either ovate to lanceolate, 30–70(–120) × 12–18(–45) mm, or 1-pinnately lobed, primary lobes 3–7, ovate to lanceolate [linear], (10–)25–80+ × (5–)10–40+ mm [blades 2(–3)-pinnatisect], bases truncate to cuneate, ultimate margins serrate or entire, usually ciliate, apices acute to attenuate, faces pilosulous to sparsely hirtellous or glabrate.

Peduncles

10–40(–80) mm.

10–20(–90) mm.

Involucres

campanulate to hemispheric or broader, (4–)5–7(–12) × (3–)6–12(–15+) mm.

turbinate to campanulate, 5–6 × (6–)7–8 mm.

Ray florets

usually 0, sometimes 1–5;

laminae orange yellowish, 4–8 mm.

0 or (3–)5–8+;

laminae whitish to pinkish [yellowish], 2–3 or 7–15+ mm.

Disc florets

(5–)20–60(–150+);

corollas pale yellow to orange, (2–)3–4 mm (gradually ampliate, anthers usually pale).

20–40(–80+);

corollas yellowish, (2–)3–5 mm.

Phyllaries

(6–)7–8(–13), elliptic-ovate to lance-ovate, (4–)6–9(–12) mm.

(7–)8–9(–13), lanceolate to oblanceolate, 4–6 mm.

Calyculi

of (2–)6–7(–10) ± spreading, oblanceolate or lanceolate to linear, ± foliaceous bractlets or bracts 7–35(–60) mm, margins (entire or serrate) sometimes sparsely ciliate, abaxial faces hispidulous near bases, distally glabrous.

(6–)7–9(–13) ± appressed, spatulate to linear bractlets (3–)4–5 mm, margins ciliate, abaxial faces usually hispidulous to puberulent.

Heads

borne singly or in 2s or 3s.

usually borne singly, sometimes in open, ± corymbiform arrays.

Cypselae

blackish to purplish or brown, ± flattened, sometimes weakly 3(–4)-angled, usually cuneate to linear, outer (3–)6–7(–10) mm, inner (4–)6–9(–11) mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces ± 1-nerved, usually smooth, seldom notably tuberculate, glabrous or sparsely strigillose;

pappi 0, or of (1–)3–3(–4+) ± erect to spreading, retrorsely barbed awns (0.2–)2–3(–6) mm.

outer red-brown, ± flat, linear to narrowly cuneate, (3–)4–5+ mm, margins antrorsely hispidulous, apices ± truncate or somewhat attenuate, faces obscurely 2-grooved, sometimes tuberculate-hispidulous;

inner blackish, ± equally 4-angled, linear-fusiform, 7–16 mm, margins antrorsely hispidulous, apices ± attenuate, faces 2-grooved, tuberculate-hispidulous to sparsely strigillose;

pappi 0, or of 2–3(–5), erect to divergent, retrorsely barbed awns (0.5–)2–4 mm.

2n

= 48.

= 24, 36, 48, 72.

Bidens tripartita

Bidens pilosa

Phenology Flowering Aug–Oct. Flowering year round.
Habitat Marshes and other wet sites Disturbed, wettish sites
Elevation 10–1700 m (0–5600 ft) 10–1900 m (0–6200 ft)
Distribution
from FNA
AK; AL; CA; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AZ; CA; CT; FL; GA; KY; LA; MA; MD; MO; MS; NC; NM; PA; SC; TX; ON; QC; Mexico; Central America [Introduced, Europe]
[WildflowerSearch map]
[BONAP county map]
Discussion

Plants with cypsela mid-nerves strongly developed (cypselae more or less strongly 4-angled and, often, tuberculate) that are treated below as Bidens connata have been included in B. tripartita, perhaps rightly so. And some botanists have included (or advocated inclusion of) B. eatonii, B. heterodoxa, and/or B. infirma in B. tripartita, as well, perhaps rightly so.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

R. Ballard (1986) adopted a narrower circumscription of Bidens pilosa than that used here. He used: B. pilosa for plants with outer phyllaries 7–10, ray florets usually 0 (when present, laminae 2–3 mm), disc florets 35–75, pappi of 3(–5) awns 1–3 mm, and 2n = 72; B. alba for plants with outer phyllaries (8–)12(–16), ray florets 5–8 (laminae 5–16 mm), pappi of 2 awns 1–2 mm, and 2n = 48; and B. odorata for plants with outer phyllaries (6–)8(–12), ray florets 5–8 (laminae 3–18 mm), disc florets 12–61, pappi 0, or of 1–2 awns 1–3 mm, and 2n = 24.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 217. FNA vol. 21, p. 211.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens
Sibling taxa
B. amplissima, B. aristosa, B. aurea, B. beckii, B. bidentoides, B. bigelovii, B. bipinnata, B. cernua, B. connata, B. discoidea, B. eatonii, B. frondosa, B. heterodoxa, B. heterosperma, B. hyperborea, B. laevis, B. lemmonii, B. leptocephala, B. mitis, B. pilosa, B. polylepis, B. tenuisecta, B. trichosperma, B. vulgata
B. amplissima, B. aristosa, B. aurea, B. beckii, B. bidentoides, B. bigelovii, B. bipinnata, B. cernua, B. connata, B. discoidea, B. eatonii, B. frondosa, B. heterodoxa, B. heterosperma, B. hyperborea, B. laevis, B. lemmonii, B. leptocephala, B. mitis, B. polylepis, B. tenuisecta, B. trichosperma, B. tripartita, B. vulgata
Synonyms B. comosa B. alba, B. alba var. radiata, B. odorata, B. pilosa var. radiata
Name authority Linnaeus: Sp. Pl. 2: 831. (1753) Linnaeus: Sp. Pl. 2: 832. (1753)
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