Bidens tripartita |
Bidens laevis |
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bident à trois divisions, three-lobed beggar-ticks, three-lobed beggars-ticks, three-part beggarticks, threelobe beggarticks, tickseed |
bur-marigold, smooth beggar-ticks, smooth beggartick |
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Habit | Annuals, (5–)20–70(–200) cm. | Annuals (sometimes persisting), (10–)20–60(–120+) cm. |
Leaves | sessile or petioles 5–15(–35+) mm (± winged); blades ± elliptic to ovate or lanceolate, 40–80(–150+) × 15–40(–60+) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices acute to acuminate, faces glabrous or hirtellous. |
(sometimes in 3s or 4s) sessile; blades obovate or elliptic to lanceolate or linear, (20–)50–100(–160+) × (5–)10–25(–40+) mm, bases cuneate to rounded, margins usually coarsely dentate to serrate, sometimes ciliate, apices acute to acuminate, faces glabrous. |
Peduncles | 10–40(–80) mm. |
(10–)20–60 mm. |
Involucres | campanulate to hemispheric or broader, (4–)5–7(–12) × (3–)6–12(–15+) mm. |
turbinate to hemispheric or broader, (4–)6–8(–10+) × 8–12+ mm. |
Ray florets | usually 0, sometimes 1–5; laminae orange yellowish, 4–8 mm. |
usually 7–8, rarely 0; laminae orange-yellow, (10–)15–25(–30) mm. |
Disc florets | (5–)20–60(–150+); corollas pale yellow to orange, (2–)3–4 mm (gradually ampliate, anthers usually pale). |
(25–)60–100(–150+); corollas yellow to orange-yellow, 3–6.5 mm. |
Phyllaries | (6–)7–8(–13), elliptic-ovate to lance-ovate, (4–)6–9(–12) mm. |
8–12, ovate or obovate to lance-oblong, (4–)6–8(–10+) mm (tips often orange to purplish, as are tips of paleae). |
Calyculi | of (2–)6–7(–10) ± spreading, oblanceolate or lanceolate to linear, ± foliaceous bractlets or bracts 7–35(–60) mm, margins (entire or serrate) sometimes sparsely ciliate, abaxial faces hispidulous near bases, distally glabrous. |
of 5–7(–9+) erect or spreading to reflexed, oblanceolate or lanceolate to linear, often foliaceous bractlets or bracts (6–)10–12(–20+) mm, margins usually ciliate, abaxial faces glabrous or bases hispidulous. |
Heads | borne singly or in 2s or 3s. |
(erect at flowering, sometimes nodding in fruit) borne singly or in open, ± corymbiform arrays. |
Cypselae | blackish to purplish or brown, ± flattened, sometimes weakly 3(–4)-angled, usually cuneate to linear, outer (3–)6–7(–10) mm, inner (4–)6–9(–11) mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces ± 1-nerved, usually smooth, seldom notably tuberculate, glabrous or sparsely strigillose; pappi 0, or of (1–)3–3(–4+) ± erect to spreading, retrorsely barbed awns (0.2–)2–3(–6) mm. |
blackish, red-brown, or stramineous, obcompressed, flattened or unequally 3–4-angled, ± cuneate, outer 6–8 mm, inner 8–10 mm, margins retrorsely ciliate or barbed, apices ± truncate to convex, faces ± 1-nerved, ± striate, glabrous; pappi of 2–4 ± erect, retrorsely barbed awns 3–5 mm. |
2n | = 48. |
= 22, 24. |
Bidens tripartita |
Bidens laevis |
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Phenology | Flowering Aug–Oct. | Flowering Aug–Oct(–Dec). |
Habitat | Marshes and other wet sites | Meadows, marshes, and margins of pools, streams, estuaries |
Elevation | 10–1700 m (0–5600 ft) | 0–2800 m (0–9200 ft) |
Distribution |
AK; AL; CA; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
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AL; AR; AZ; CA; CO; CT; DC; DE; FL; GA; IN; KY; LA; MA; MD; ME; MO; MS; NC; NH; NJ; NM; NV; NY; OH; PA; RI; SC; TN; TX; VA; WV; Mexico; Central America; South America [Introduced Pacific Islands (Hawaii)]
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Discussion | Plants with cypsela mid-nerves strongly developed (cypselae more or less strongly 4-angled and, often, tuberculate) that are treated below as Bidens connata have been included in B. tripartita, perhaps rightly so. And some botanists have included (or advocated inclusion of) B. eatonii, B. heterodoxa, and/or B. infirma in B. tripartita, as well, perhaps rightly so. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Bidens laevis and B. cernua are similar in gross appearance; specimens of one are sometimes misidentified as the other. Perhaps they represent extremes of a single species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 217. | FNA vol. 21, p. 215. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens |
Sibling taxa | ||
Synonyms | B. comosa | Helianthus laevis, B. chrysanthemoides, B. helianthoides, B. nashii, B. quadriaristata |
Name authority | Linnaeus: Sp. Pl. 2: 831. (1753) | (Linnaeus) Britton: Prelim. Cat., 29. (1888) |
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