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bident à trois divisions, three-lobed beggar-ticks, three-lobed beggars-ticks, three-part beggarticks, threelobe beggarticks, tickseed

Rocky Mountain beggarticks

Habit Annuals, (5–)20–70(–200) cm. Annuals, (5–)10–30(–60+) cm.
Leaves

sessile or petioles 5–15(–35+) mm (± winged);

blades ± elliptic to ovate or lanceolate, 40–80(–150+) × 15–40(–60+) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices acute to acuminate, faces glabrous or hirtellous.

petioles 3–20 mm;

blades usually rounded-deltate overall, 10–30+ × 5–20+ mm, 1–2-pinnatisect, ultimate lobes linear to filiform, 2–25 × 0.5–1(–2) mm, bases obscurely cuneate, ultimate margins minutely, if at all, ciliate, apices obtuse to acute (sometimes apiculate), faces glabrous or sparsely hirtellous.

Peduncles

10–40(–80) mm.

10–50(–100) mm.

Involucres

campanulate to hemispheric or broader, (4–)5–7(–12) × (3–)6–12(–15+) mm.

± cylindric, 3.5–5 × (1–)2–3 mm.

Ray florets

usually 0, sometimes 1–5;

laminae orange yellowish, 4–8 mm.

0 or 1–3+;

laminae yellowish, 1–4 mm.

Disc florets

(5–)20–60(–150+);

corollas pale yellow to orange, (2–)3–4 mm (gradually ampliate, anthers usually pale).

5–9(–13);

corollas yellowish, 1–1.5 mm.

Phyllaries

(6–)7–8(–13), elliptic-ovate to lance-ovate, (4–)6–9(–12) mm.

5–6, lance-elliptic to lanceolate, 2.5–4(–5) mm.

Calyculi

of (2–)6–7(–10) ± spreading, oblanceolate or lanceolate to linear, ± foliaceous bractlets or bracts 7–35(–60) mm, margins (entire or serrate) sometimes sparsely ciliate, abaxial faces hispidulous near bases, distally glabrous.

of (1–)3(–5), appressed to spreading, linear bractlets (1–)3(–4) mm, margins minutely, if at all, ciliate, abaxial faces usually glabrous, sometimes sparsely hispidulous.

Heads

borne singly or in 2s or 3s.

usually borne singly.

Cypselae

blackish to purplish or brown, ± flattened, sometimes weakly 3(–4)-angled, usually cuneate to linear, outer (3–)6–7(–10) mm, inner (4–)6–9(–11) mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces ± 1-nerved, usually smooth, seldom notably tuberculate, glabrous or sparsely strigillose;

pappi 0, or of (1–)3–3(–4+) ± erect to spreading, retrorsely barbed awns (0.2–)2–3(–6) mm.

red-brown to blackish (sometimes with lighter blotches), outer ± obcompressed and linear, 3–5 mm, margins not ciliate, apices ± truncate, faces 2-grooved, usually glabrous;

inner similar, ± equally 4-angled, linear-fusiform, (6–)7–12 mm, apices ± attenuate;

pappi 0, or of 2(–3) spreading to divergent, retrorsely barbed awns 1–2 mm.

2n

= 48.

Bidens tripartita

Bidens heterosperma

Phenology Flowering Aug–Oct. Flowering Sep–Oct.
Habitat Marshes and other wet sites Seeps on rocky slopes
Elevation 10–1700 m [30–5600 ft] 1500–2800 m [4900–9200 ft]
Distribution
from FNA
AK; AL; CA; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; Mexico (Baja California, Chihuahua, Sinaloa, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Plants with cypsela mid-nerves strongly developed (cypselae more or less strongly 4-angled and, often, tuberculate) that are treated below as Bidens connata have been included in B. tripartita, perhaps rightly so. And some botanists have included (or advocated inclusion of) B. eatonii, B. heterodoxa, and/or B. infirma in B. tripartita, as well, perhaps rightly so.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 217. Treatment authors: John L. Strother, Ronald R. Weedon. FNA vol. 21, p. 209. Treatment authors: John L. Strother, Ronald R. Weedon.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens
Sibling taxa
B. amplissima, B. aristosa, B. aurea, B. beckii, B. bidentoides, B. bigelovii, B. bipinnata, B. cernua, B. connata, B. discoidea, B. eatonii, B. frondosa, B. heterodoxa, B. heterosperma, B. hyperborea, B. laevis, B. lemmonii, B. leptocephala, B. mitis, B. pilosa, B. polylepis, B. tenuisecta, B. trichosperma, B. vulgata
B. amplissima, B. aristosa, B. aurea, B. beckii, B. bidentoides, B. bigelovii, B. bipinnata, B. cernua, B. connata, B. discoidea, B. eatonii, B. frondosa, B. heterodoxa, B. hyperborea, B. laevis, B. lemmonii, B. leptocephala, B. mitis, B. pilosa, B. polylepis, B. tenuisecta, B. trichosperma, B. tripartita, B. vulgata
Synonyms B. comosa
Name authority Linnaeus: Sp. Pl. 2: 831. (1753) A. Gray: Smithsonian Contr. Knowl. 5(6): 90. (1853)
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