Bidens pilosa
Bidens heterodoxa
bident poilu, cobbler's pegs, common beggar-ticks, hairy beggar-ticks, Spanish needle
bident différent, connecticut beggarticks
petioles 10–30(–70) mm;
blades either ovate to lanceolate, 30–70(–120) × 12–18(–45) mm, or 1-pinnately lobed, primary lobes 3–7, ovate to lanceolate [linear], (10–)25–80+ × (5–)10–40+ mm [blades 2(–3)-pinnatisect], bases truncate to cuneate, ultimate margins serrate or entire, usually ciliate, apices acute to attenuate, faces pilosulous to sparsely hirtellous or glabrate.
petioles (narrowly, if at all, winged) 5–20(–35+) mm;
blades lanceolate to lance-linear, (20–)30–50(–80+) × (3–)6–20(–30+) mm, sometimes laciniately 1-pinnatisect with 2–4+ lobes near bases, bases cuneate, margins incised to serrate, seldom ciliate, apices acute to attenuate, faces glabrous.
10–20(–90) mm.
10–35(–55) mm.
turbinate to campanulate, 5–6 × (6–)7–8 mm.
usually obconic to broadly campanulate, rarely hemispheric, (4–)6–7 × (4–)5–7 mm.
0 or (3–)5–8+;
laminae whitish to pinkish [yellowish], 2–3 or 7–15+ mm.
usually 0, sometimes 1–3+;
laminae orange yellowish, 2–5(–10) mm.
20–40(–80+);
corollas yellowish, (2–)3–5 mm.
(7–)20–70+;
corollas yellowish, 2.5–3.5 mm.
(7–)8–9(–13), lanceolate to oblanceolate, 4–6 mm.
(4–)5–8,oblong to lance-ovate, (4–)5–8+ mm.
(6–)7–9(–13) ± appressed, spatulate to linear bractlets (3–)4–5 mm, margins ciliate, abaxial faces usually hispidulous to puberulent.
of (2–)3–5(–7) ± erect, oblanceolate or spatulate to linear, ± foliaceous bractlets or bracts (6–)10–25(–40+) mm, margins rarely ciliate, abaxial faces usually glabrous, sometimes sparsely hispidulous.
usually borne singly, sometimes in open, ± corymbiform arrays.
borne singly or in 2s or 3s.
outer red-brown, ± flat, linear to narrowly cuneate, (3–)4–5+ mm, margins antrorsely hispidulous, apices ± truncate or somewhat attenuate, faces obscurely 2-grooved, sometimes tuberculate-hispidulous;
inner blackish, ± equally 4-angled, linear-fusiform, 7–16 mm, margins antrorsely hispidulous, apices ± attenuate, faces 2-grooved, tuberculate-hispidulous to sparsely strigillose;
pappi 0, or of 2–3(–5), erect to divergent, retrorsely barbed awns (0.5–)2–4 mm.
outer red-brown, ± flattened, sometimes 3–4-angled, broadly cuneate, 4–5(–6) mm, inner blackish, narrowly cuneate, (5–)6–7(–8+) mm, margins glabrous or sparsely antrorsely strigillose,apices ± truncate to convex, faces ± 1-nerved, glabrous or sparsely setulose, usually smooth, sometimes ± tuberculate;
pappi 0, or of 2(–4) spreading to divergent, antrorsely or retrorsely barbed awns (0.1–)1–3.5+ mm.
= 24, 36, 48, 72.
= 48.
Bidens pilosa
Bidens heterodoxa
R. Ballard (1986) adopted a narrower circumscription of Bidens pilosa than that used here. He used: B. pilosa for plants with outer phyllaries 7–10, ray florets usually 0 (when present, laminae 2–3 mm), disc florets 35–75, pappi of 3(–5) awns 1–3 mm, and 2n = 72; B. alba for plants with outer phyllaries (8–)12(–16), ray florets 5–8 (laminae 5–16 mm), pappi of 2 awns 1–2 mm, and 2n = 48; and B. odorata for plants with outer phyllaries (6–)8(–12), ray florets 5–8 (laminae 3–18 mm), disc florets 12–61, pappi 0, or of 1–2 awns 1–3 mm, and 2n = 24.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Bidens heterodoxa may be better treated as part of B. tripartita.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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