Bidens connata |
Bidens tripartita |
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bident conné, purple-stem beggar-ticks, purplestem beggars-ticks, swamp beggar-ticks |
bident à trois divisions, three-lobed beggar-ticks, three-lobed beggars-ticks, three-part beggarticks, threelobe beggarticks, tickseed |
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Habit | Annuals, (5–)25–150(–200) cm. | Annuals, (5–)20–70(–200) cm. |
Leaves | sessile or petioles (± winged) 5–15(–35+) mm; blades ± elliptic to lanceolate, (20–)40–100(–200+) × 10–30(–70) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices attenuate, faces glabrous or hirtellous. |
sessile or petioles 5–15(–35+) mm (± winged); blades ± elliptic to ovate or lanceolate, 40–80(–150+) × 15–40(–60+) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices acute to acuminate, faces glabrous or hirtellous. |
Peduncles | 10–35(–80) mm. |
10–40(–80) mm. |
Involucres | campanulate to hemispheric or broader, (4–)6–12 × (3–)6–9(–15+) mm. |
campanulate to hemispheric or broader, (4–)5–7(–12) × (3–)6–12(–15+) mm. |
Ray florets | usually 0, sometimes 1–5+; laminae yellowish, 4–8 mm. |
usually 0, sometimes 1–5; laminae orange yellowish, 4–8 mm. |
Disc florets | (5–)20–40(–60+); corollas pale yellow to orange, 2–3 mm (± abruptly ampliate, anthers usually dark, blackish). |
(5–)20–60(–150+); corollas pale yellow to orange, (2–)3–4 mm (gradually ampliate, anthers usually pale). |
Phyllaries | (6–)8(–9), elliptic to oblong or ovate, (4–)5–6(–12) mm. |
(6–)7–8(–13), elliptic-ovate to lance-ovate, (4–)6–9(–12) mm. |
Calyculi | of 2–6+ spreading or ascending, lanceolate, linear, oblanceolate, oblong, or spatulate, ± foliaceous bractlets or bracts 10–25(–35+) mm, margins (entire or serrate) usually ciliate, abaxial faces hispidulous near bases, distally glabrous. |
of (2–)6–7(–10) ± spreading, oblanceolate or lanceolate to linear, ± foliaceous bractlets or bracts 7–35(–60) mm, margins (entire or serrate) sometimes sparsely ciliate, abaxial faces hispidulous near bases, distally glabrous. |
Heads | borne singly or in 2s or 3s. |
borne singly or in 2s or 3s. |
Cypselae | blackish to purplish or brown, usually (at least inner) ± 4-angled, usually cuneate to linear, sometimes ± obpyramidal, outer (3–)4–7 mm, inner (4–)5–8 mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces usually strongly 1-nerved, usually tuberculate, glabrous or setulose; pappi 0, or of (1–)2–4(–6), ± erect to spreading, antrorsely or retrorsely barbed awns (0.2–)2–5 mm. |
blackish to purplish or brown, ± flattened, sometimes weakly 3(–4)-angled, usually cuneate to linear, outer (3–)6–7(–10) mm, inner (4–)6–9(–11) mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces ± 1-nerved, usually smooth, seldom notably tuberculate, glabrous or sparsely strigillose; pappi 0, or of (1–)3–3(–4+) ± erect to spreading, retrorsely barbed awns (0.2–)2–3(–6) mm. |
2n | = 48. |
= 48. |
Bidens connata |
Bidens tripartita |
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Phenology | Flowering Aug–Oct. | Flowering Aug–Oct. |
Habitat | Marshes and other wet sites | Marshes and other wet sites |
Elevation | 10–1700 m (0–5600 ft) | 10–1700 m (0–5600 ft) |
Distribution |
CT; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; SD; TN; VA; WI; WV; NB; NF; NS; ON; PE; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
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AK; AL; CA; CO; CT; DC; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NM; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; UT; VA; VT; WA; WI; WV; WY; AB; BC; MB; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
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Discussion | Bidens connata may be better treated as part of B. tripartita. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants with cypsela mid-nerves strongly developed (cypselae more or less strongly 4-angled and, often, tuberculate) that are treated below as Bidens connata have been included in B. tripartita, perhaps rightly so. And some botanists have included (or advocated inclusion of) B. eatonii, B. heterodoxa, and/or B. infirma in B. tripartita, as well, perhaps rightly so. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 217. | FNA vol. 21, p. 217. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens |
Sibling taxa | ||
Synonyms | B. connata var. ambiversa, B. connata var. anomala, B. connata var. fallax, B. connata var. gracilipes, B. connata var. inundata, B. connata var. petiolata, B. connata var. pinnata, B. connata var. submutica | B. comosa |
Name authority | Muhlenberg ex Willdenow: Sp. Pl. 3: 1718. (1803) | Linnaeus: Sp. Pl. 2: 831. (1753) |
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