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bident conné, purple-stem beggar-ticks, purplestem beggars-ticks, swamp beggar-ticks

bident différent, connecticut beggarticks

Habit Annuals, (5–)25–150(–200) cm. Annuals, (5–)25–35(–55+) cm.
Leaves

sessile or petioles (± winged) 5–15(–35+) mm;

blades ± elliptic to lanceolate, (20–)40–100(–200+) × 10–30(–70) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices attenuate, faces glabrous or hirtellous.

petioles (narrowly, if at all, winged) 5–20(–35+) mm;

blades lanceolate to lance-linear, (20–)30–50(–80+) × (3–)6–20(–30+) mm, sometimes laciniately 1-pinnatisect with 2–4+ lobes near bases, bases cuneate, margins incised to serrate, seldom ciliate, apices acute to attenuate, faces glabrous.

Peduncles

10–35(–80) mm.

10–35(–55) mm.

Involucres

campanulate to hemispheric or broader, (4–)6–12 × (3–)6–9(–15+) mm.

usually obconic to broadly campanulate, rarely hemispheric, (4–)6–7 × (4–)5–7 mm.

Ray florets

usually 0, sometimes 1–5+;

laminae yellowish, 4–8 mm.

usually 0, sometimes 1–3+;

laminae orange yellowish, 2–5(–10) mm.

Disc florets

(5–)20–40(–60+);

corollas pale yellow to orange, 2–3 mm (± abruptly ampliate, anthers usually dark, blackish).

(7–)20–70+;

corollas yellowish, 2.5–3.5 mm.

Phyllaries

(6–)8(–9), elliptic to oblong or ovate, (4–)5–6(–12) mm.

(4–)5–8,oblong to lance-ovate, (4–)5–8+ mm.

Calyculi

of 2–6+ spreading or ascending, lanceolate, linear, oblanceolate, oblong, or spatulate, ± foliaceous bractlets or bracts 10–25(–35+) mm, margins (entire or serrate) usually ciliate, abaxial faces hispidulous near bases, distally glabrous.

of (2–)3–5(–7) ± erect, oblanceolate or spatulate to linear, ± foliaceous bractlets or bracts (6–)10–25(–40+) mm, margins rarely ciliate, abaxial faces usually glabrous, sometimes sparsely hispidulous.

Heads

borne singly or in 2s or 3s.

borne singly or in 2s or 3s.

Cypselae

blackish to purplish or brown, usually (at least inner) ± 4-angled, usually cuneate to linear, sometimes ± obpyramidal, outer (3–)4–7 mm, inner (4–)5–8 mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces usually strongly 1-nerved, usually tuberculate, glabrous or setulose;

pappi 0, or of (1–)2–4(–6), ± erect to spreading, antrorsely or retrorsely barbed awns (0.2–)2–5 mm.

outer red-brown, ± flattened, sometimes 3–4-angled, broadly cuneate, 4–5(–6) mm, inner blackish, narrowly cuneate, (5–)6–7(–8+) mm, margins glabrous or sparsely antrorsely strigillose,apices ± truncate to convex, faces ± 1-nerved, glabrous or sparsely setulose, usually smooth, sometimes ± tuberculate;

pappi 0, or of 2(–4) spreading to divergent, antrorsely or retrorsely barbed awns (0.1–)1–3.5+ mm.

2n

= 48.

= 48.

Bidens connata

Bidens heterodoxa

Phenology Flowering Aug–Oct. Flowering Aug–Oct.
Habitat Marshes and other wet sites Borders of estuaries, lakes, marshes, streams
Elevation 10–1700 m (0–5600 ft) 0–10 m (0–0 ft)
Distribution
from FNA
CT; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; SD; TN; VA; WI; WV; NB; NF; NS; ON; PE; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
[WildflowerSearch map]
from FNA
CT; NB; PE; QC
[BONAP county map]
Discussion

Bidens connata may be better treated as part of B. tripartita.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Bidens heterodoxa may be better treated as part of B. tripartita.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 217. FNA vol. 21, p. 218.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens
Sibling taxa
B. amplissima, B. aristosa, B. aurea, B. beckii, B. bidentoides, B. bigelovii, B. bipinnata, B. cernua, B. discoidea, B. eatonii, B. frondosa, B. heterodoxa, B. heterosperma, B. hyperborea, B. laevis, B. lemmonii, B. leptocephala, B. mitis, B. pilosa, B. polylepis, B. tenuisecta, B. trichosperma, B. tripartita, B. vulgata
B. amplissima, B. aristosa, B. aurea, B. beckii, B. bidentoides, B. bigelovii, B. bipinnata, B. cernua, B. connata, B. discoidea, B. eatonii, B. frondosa, B. heterosperma, B. hyperborea, B. laevis, B. lemmonii, B. leptocephala, B. mitis, B. pilosa, B. polylepis, B. tenuisecta, B. trichosperma, B. tripartita, B. vulgata
Synonyms B. connata var. ambiversa, B. connata var. anomala, B. connata var. fallax, B. connata var. gracilipes, B. connata var. inundata, B. connata var. petiolata, B. connata var. pinnata, B. connata var. submutica B. tripartita var. heterodoxa, B. heterodoxa var. agnostica, B. heterodoxa var. monardifolia, B. heterodoxa var. orthodoxa, B. infirma
Name authority Muhlenberg ex Willdenow: Sp. Pl. 3: 1718. (1803) (Fernald) Fernald & H. St. John: Rhodora 17: 23. (1915)
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