Bidens connata |
Bidens bipinnata |
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bident conné, purple-stem beggar-ticks, purplestem beggars-ticks, swamp beggar-ticks |
Spanish needles, Spanish needles beggar-ticks |
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Habit | Annuals, (5–)25–150(–200) cm. | Annuals, (15–)30–100(–150+) cm. |
Leaves | sessile or petioles (± winged) 5–15(–35+) mm; blades ± elliptic to lanceolate, (20–)40–100(–200+) × 10–30(–70) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices attenuate, faces glabrous or hirtellous. |
petioles 20–50 mm; blades rounded-deltate to ovate or lanceolate overall, (20–)30–70+ × (20–)30–60+ mm, (1–)2(–3)-pinnatisect, ultimate lobes obovate or lanceolate, 15–45+ × 10–25+ mm, bases truncate to cuneate, ultimate margins entire, sometimes ciliolate, apices rounded to acute or attenuate, faces usually glabrous, sometimes hirtellous. |
Peduncles | 10–35(–80) mm. |
(10–)20–50(–100) mm. |
Involucres | campanulate to hemispheric or broader, (4–)6–12 × (3–)6–9(–15+) mm. |
± campanulate, 5–7 × 3–4(–5) mm. |
Ray florets | usually 0, sometimes 1–5+; laminae yellowish, 4–8 mm. |
0 or 3–5+; laminae yellowish or whitish, 1–2(–3) mm. |
Disc florets | (5–)20–40(–60+); corollas pale yellow to orange, 2–3 mm (± abruptly ampliate, anthers usually dark, blackish). |
10–20(–30+); corollas yellowish to whitish, 2–3 mm. |
Phyllaries | (6–)8(–9), elliptic to oblong or ovate, (4–)5–6(–12) mm. |
8–12, lanceolate to linear, 4–6 mm. |
Calyculi | of 2–6+ spreading or ascending, lanceolate, linear, oblanceolate, oblong, or spatulate, ± foliaceous bractlets or bracts 10–25(–35+) mm, margins (entire or serrate) usually ciliate, abaxial faces hispidulous near bases, distally glabrous. |
of (7–)8(–10) linear bractlets 3–5 mm, ± appressed, margins ciliate, abaxial faces usually glabrous. |
Heads | borne singly or in 2s or 3s. |
usually borne singly, sometimes in ± corymbiform arrays. |
Cypselae | blackish to purplish or brown, usually (at least inner) ± 4-angled, usually cuneate to linear, sometimes ± obpyramidal, outer (3–)4–7 mm, inner (4–)5–8 mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces usually strongly 1-nerved, usually tuberculate, glabrous or setulose; pappi 0, or of (1–)2–4(–6), ± erect to spreading, antrorsely or retrorsely barbed awns (0.2–)2–5 mm. |
red-brown, outer weakly obcompressed, 7–15 mm, inner ± 4-angled, linear to linear-fusiform, 12–18 mm, margins not ciliate, apices ± attenuate, faces 2-grooved, often tuberculate-hispidulous; pappi of (2–)3–4, erect to divergent, retrorsely barbed awns 2–4 mm. |
2n | = 48. |
= 24, 72. |
Bidens connata |
Bidens bipinnata |
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Phenology | Flowering Aug–Oct. | Flowering (Jun–)Aug–Sep(–Oct). |
Habitat | Marshes and other wet sites | Fields, forests, disturbed, wettish sites |
Elevation | 10–1700 m (0–5600 ft) | 10–1800 m (0–5900 ft) |
Distribution |
CT; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; SD; TN; VA; WI; WV; NB; NF; NS; ON; PE; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
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AL; AR; AZ; CT; DC; FL; GA; IA; IL; IN; KS; KY; LA; MD; MO; MS; NC; NE; NJ; NM; NY; OH; OK; PA; RI; SC; TN; TX; VA; WV; NB; ON; South America; Europe; Asia; Pacific Islands
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Discussion | Bidens connata may be better treated as part of B. tripartita. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Bidens bipinnata is probably native in eastern Asia and introduced in South America, Europe, Asia, and Pacific Islands. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 217. | FNA vol. 21, p. 210. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens |
Sibling taxa | ||
Synonyms | B. connata var. ambiversa, B. connata var. anomala, B. connata var. fallax, B. connata var. gracilipes, B. connata var. inundata, B. connata var. petiolata, B. connata var. pinnata, B. connata var. submutica | B. bipinnata var. biternatoides |
Name authority | Muhlenberg ex Willdenow: Sp. Pl. 3: 1718. (1803) | Linnaeus: Sp. Pl. 2: 832. (1753) |
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