Bidens connata |
Asteraceae tribe Heliantheae subtribe Coreopsidinae |
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bident conné, purple-stem beggar-ticks, purplestem beggars-ticks, swamp beggar-ticks |
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Habit | Annuals, (5–)25–150(–200) cm. | Annuals, perennials, subshrubs, or shrubs, 10–400 cm (sometimes rhizomatous or with cormiform bases, stoloniferous in Coreopsis auriculata). | ||||||||||||||||||||||||||||
Leaves | sessile or petioles (± winged) 5–15(–35+) mm; blades ± elliptic to lanceolate, (20–)40–100(–200+) × 10–30(–70) mm, sometimes laciniately 1-pinnatisect with 1–4+ lobes near bases, bases cuneate, margins entire or dentate to serrate, usually ciliate, apices attenuate, faces glabrous or hirtellous. |
usually cauline (sometimes mostly basal); usually mostly opposite (distal sometimes alternate), rarely whorled; petiolate or sessile; blades (often pinnately or palmately lobed, sometimes compound), or lobes or leaflets, mostly deltate, elliptic, filiform, lanceolate, linear, oblanceolate, or ovate, ultimate margins entire or toothed, faces usually glabrous or glabrate, sometimes hairy (rarely, if ever, gland-dotted). |
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Peduncles | 10–35(–80) mm. |
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Involucres | campanulate to hemispheric or broader, (4–)6–12 × (3–)6–9(–15+) mm. |
mostly campanulate, cylindric, turbinate, or hemispheric or broader. |
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Receptacles | mostly flat to convex, paleate (paleae often stramineous to white with red-brown to purplish striae, orbiculate to oblong or linear, ± flat to slightly cupped). |
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Ray florets | usually 0, sometimes 1–5+; laminae yellowish, 4–8 mm. |
0 or 1–21+ (more in cultivars), pistillate and fertile, or neuter, or styliferous and sterile; corollas usually yellow to orange (sometimes bicolored: yellow to orange with brown, red-brown, or purple), sometimes cyanic or white. |
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Disc florets | (5–)20–40(–60+); corollas pale yellow to orange, 2–3 mm (± abruptly ampliate, anthers usually dark, blackish). |
3–150+, usually bisexual, fertile (functionally staminate in Dicranocarpus); corollas usually yellow to orange, sometimes brown, red-brown, or purple, tubes usually shorter than, sometimes equaling funnelform throats (longer than throats in some Thelesperma spp.), lobes (3–)5, ± deltate to lance-ovate; (staminal filaments hairy in Cosmos) anther thecae pale or dark; stigmatic papillae in 2 lines. |
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Phyllaries | (6–)8(–9), elliptic to oblong or ovate, (4–)5–6(–12) mm. |
persistent, 3–34+ in ± 2 series (usually distinct, notably connate in Thelesperma), usually ± membranous, margins usually ± scarious. |
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Calyculi | of 2–6+ spreading or ascending, lanceolate, linear, oblanceolate, oblong, or spatulate, ± foliaceous bractlets or bracts 10–25(–35+) mm, margins (entire or serrate) usually ciliate, abaxial faces hispidulous near bases, distally glabrous. |
usually of 3–8(–21+) bractlets or bracts (usually ± herbaceous, usually shorter than phyllaries and/or reflexed, sometimes ± foliaceous and surpassing phyllaries). |
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Heads | borne singly or in 2s or 3s. |
radiate or discoid, borne singly or in ± corymbiform, cymiform, or paniculiform arrays. |
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Cypselae | blackish to purplish or brown, usually (at least inner) ± 4-angled, usually cuneate to linear, sometimes ± obpyramidal, outer (3–)4–7 mm, inner (4–)5–8 mm, margins proximally antrorsely to patently, distally retrorsely, barbed, apices ± truncate to concave, faces usually strongly 1-nerved, usually tuberculate, glabrous or setulose; pappi 0, or of (1–)2–4(–6), ± erect to spreading, antrorsely or retrorsely barbed awns (0.2–)2–5 mm. |
usually either obcompressed to obflattened, usually cuneate, linear, oblong, orbiculate, or ovate (bodies unequally 3–4-angled, margins often winged), or (all or inner) ± equally 4-angled and linear-fusiform, sometimes ± beaked, faces glabrous or hairy (often striate, tuberculate, or papillate); pappi 0, or persistent, of (1–)2–4(–8), usually ± barbellate awns or scales, rarely coroniform or of 1–2, smooth to ciliate or barbed awns or scales. |
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2n | = 48. |
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Bidens connata |
Asteraceae tribe Heliantheae subtribe Coreopsidinae |
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Phenology | Flowering Aug–Oct. | |||||||||||||||||||||||||||||
Habitat | Marshes and other wet sites | |||||||||||||||||||||||||||||
Elevation | 10–1700 m (0–5600 ft) | |||||||||||||||||||||||||||||
Distribution |
CT; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; PA; RI; SC; SD; TN; VA; WI; WV; NB; NF; NS; ON; PE; QC; Europe; Asia; n Africa [Introduced in Pacific Islands, Australia]
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Mostly subtropical and warm-temperate New World and Old World |
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Discussion | Bidens connata may be better treated as part of B. tripartita. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 20, species 374 (7 genera, 69 species in the flora). The circumscription of Coreopsidinae followed here (H. Robinson 1981) is a bit narrower than the traditional one. The subtribe is remarkable among Heliantheae for having a relatively high number of species native in subtropical and tropical Old World, especially Africa. Distinctions among some genera (e.g., Bidens, Coreopsis, and Cosmos) are often subtle. In Coreopsidinae, each involucre is subtended by a calyculus of more or less herbaceous (sometimes leaflike) bractlets or bracts (sometimes surpassing the phyllaries). In keys and descriptions here, shapes, heights, and diameters given for involucres are based on the phyllaries collectively (exclusive of calyculi) at flowering; the involucres are sometimes notably larger in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 217. | FNA vol. 21, p. 183. | ||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Bidens | Asteraceae > tribe Heliantheae | ||||||||||||||||||||||||||||
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Synonyms | B. connata var. ambiversa, B. connata var. anomala, B. connata var. fallax, B. connata var. gracilipes, B. connata var. inundata, B. connata var. petiolata, B. connata var. pinnata, B. connata var. submutica | subtribe Coreopsideae, subtribe Petrobiinae | ||||||||||||||||||||||||||||
Name authority | Muhlenberg ex Willdenow: Sp. Pl. 3: 1718. (1803) | Lessing: Linnaea 5: 153. (1830) | ||||||||||||||||||||||||||||
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