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California barberry, California Oregon-grape, shiny leaf mahonia

Harrison's barberry, kofa mountain barberry

Habit Shrubs, evergreen, 0.3-1.6(-7) m. Stems usually monomorphic, seldom with short axillary shoots. Shrubs, evergreen, 0.5-1.5 m. Stems often ± dimorphic, with elongate primary and somewhat elongate axillary shoots.
Bark

of 2d-year stems grayish brown, glabrous.

of 2d-year stems brown or gray, glabrous.

Leaves

(3-)5-13-foliolate;

petioles 0.5-4.5(-7.5) cm.

3-foliolate;

petioles 1.5-5 cm.

Leaflet

blades thin and ± rigid or flexible;

surfaces abaxially glossy, smooth, adaxially glossy, green;

terminal leaflet stalked, blade 2.6-6.2 × 2-4.5 cm, 1.3-1.9 times as long as wide;

lateral leaflet blades elliptic to ovate or broadly lanceolate, 1(-3)-veined from base, base broadly obtuse, truncate, or weakly cordate, margins plane to crispate, toothed, each with 5-22 teeth 0-2 mm tipped with spines to 1-3 × 0.1-0.3 mm, apex acute to rounded-obtuse.

blades thick and rigid;

surfaces abaxially ± dull, papillose, adaxially dull, rarely glossy, somewhat glaucous;

terminal leaflet sessile, blade 2.9-5.4 × 2.2-3.2 cm, 1.3-2.4 times as long as wide;

lateral leaflet blades ovate or rhombic to lanceolate, 1-3-veined from base, base acute to rounded-obtuse, margins plane or undulate, lobed, with 1-2 teeth 5-13 mm high tipped with spines to 2-3.4 × 0.3-0.4 mm, apex acuminate.

Spines

absent.

absent.

Inflorescences

racemose, dense, 25-50-flowered, 2-9 cm;

bracteoles membranous, apex rounded to broadly acute, sometimes apiculate.

racemose, rather dense, 6-11-flowered, 1.5-2.5 cm;

bracteoles membranous, apex acute or obtuse.

Flowers

anther filaments with distal pair of recurved lateral teeth.

Berries

blue, glaucous, oblong-ovoid to subspheric, 6-7 mm, juicy, solid.

blue-black, glaucous, spheric to short-ovoid, 5-6 mm, juicy, solid.

Bud

scales 3-7 mm, deciduous.

scales 1.5-3 mm, deciduous.

Anther

filaments with distal pair of recurved lateral teeth.

Berberis pinnata

Berberis harrisoniana

Phenology Flowering winter (Jan–Mar).
Habitat Shady spots in rocky canyons
Elevation 800-1100 m (2600-3600 ft)
Distribution
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
from FNA
AZ
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Berberis pinnata is very similar to B. aquifolium, and the two are sometimes difficult to separate. Some authors have used the spacing of the lateral leaflets (said to be contiguous or imbricate in B. pinnata and remote in B. aquifolium) to separate them, but the leaflets are often remote in both species and may be contiguous in B. aquifolium.

Berberis pinnata is resistant to infection by Puccinia graminis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Berberis harrisoniana is endemic to the Kofa and Ajo mountains. It has not been tested for resistance to infection by Puccinia graminis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Margins of leaf blade undulate or crispate, marginal spines to 1.2-3 × 0.2-0.3 mm; shrubs 0.3-1.6 m, self-supporting.
subsp. pinnata
1. Margins of leaf blade plane or weakly undulate, marginal spines to 1-1.6 × 0.1- 0.2 mm; shrubs 1-7 m, self-supporting or clambering over surrounding vegetation.
subsp. insularis
Source FNA vol. 3. FNA vol. 3.
Parent taxa Berberidaceae > Berberis Berberidaceae > Berberis
Sibling taxa
B. amplectens, B. aquifolium, B. bealei, B. canadensis, B. darwinii, B. dictyota, B. fendleri, B. fremontii, B. haematocarpa, B. harrisoniana, B. higginsiae, B. nervosa, B. nevinii, B. piperiana, B. pumila, B. repens, B. swaseyi, B. thunbergii, B. trifoliolata, B. vulgaris, B. wilcoxii
B. amplectens, B. aquifolium, B. bealei, B. canadensis, B. darwinii, B. dictyota, B. fendleri, B. fremontii, B. haematocarpa, B. higginsiae, B. nervosa, B. nevinii, B. pinnata, B. piperiana, B. pumila, B. repens, B. swaseyi, B. thunbergii, B. trifoliolata, B. vulgaris, B. wilcoxii
Subordinate taxa
B. pinnata subsp. insularis, B. pinnata subsp. pinnata
Synonyms Mahonia pinnata
Name authority Lagasca: Elench. Pl., 14. (1816) Kearney & Peebles: J. Wash. Acad. Sci. 29: 477. (1939)
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