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bartramia moss, rigid apple moss

Haller's apple-moss, Haller's bartramia moss

Habit Plants in dense tufts, glaucous to brownish green. Plants in lax to dense tufts, greenish to yellowish brown.
Stems

1–3 cm.

15 cm.

Leaves

stiffly erect-appressed when dry, erect-spreading when moist, narrowly lanceolate, 2.5–4 mm;

base not sheathing, shoulders absent;

margins recurved from near base to distal acumen, serrulate distally, teeth single;

apex narrowly aristate;

costa excurrent, abaxially prominent, distal abaxial surface rough;

basal laminal cell walls thin;

medial and distal cells 12–20 × 4–6 µm, prorulae low.

crisped when dry, erect-spreading or sometimes secund when moist, narrowly lanceolate to linear, 4–7 mm;

base laxly sheathing, shoulders well developed, firm;

margins revolute from shoulders to high in acumen, denticulate in shoulders, spinose-dentate distally, teeth paired;

apex long-subulate;

costa excurrent, prominent, distal abaxial surface rough;

basal laminal cell walls thin;

medial and distal cells 5–12 × 4(–10) µm, prorulae high.

Seta

0.1–1.5 cm, straight.

0.2(–0.8) cm, curved.

Sexual condition

synoicous.

autoicous or synoicous.

Capsule

erect, subglobose to ovoid, symmetric, 1.4–2 mm;

operculum conic convex;

peristome single;

exostome teeth 250–300 µm, striolate papillose proximally, smooth distally;

endostome absent.

inclined, subglobose to pyriform, asymmetric, 1.5(–2.5) mm;

operculum conic convex;

peristome double;

exostome teeth 500–550 µm, granulose proximally, papillose distally;

endostome basal membrane high, segments slightly shorter than teeth, striate papillose, cilia rudimentary.

Spores

22–26 µm.

15–24 µm.

Bartramia stricta

Bartramia halleriana

Phenology Capsules mature Jan–Aug. Spores 15-24 µm, mature Jun–Sep.
Habitat Soil, rock Crevices of shaded cliffs, rock outcrops in humid forests
Elevation low to high elevations (10-2300 m) (low to high elevations (0-7500 ft)) moderate elevations (200-500 m) (moderate elevations (700-1600 ft))
Distribution
from FNA
CA; NM; TX; BC; Europe; sw Asia (Syria, Turkey); n Africa; c Africa; Australia
[WildflowerSearch map]
from FNA
AB; BC; s South America; Europe; Asia; Pacific Islands (Hawaii, New Zealand); Australia (Tasmania)
Discussion

Bartramia stricta is a widespread Old World species occurring infrequently in the southwestern portion of the flora area, with an isolated occurrence in western Canada. The strict, appressed leaves without shoulders and with relatively uniform areolation from base to acumen aid in recognition. When fertile, the erect, symmetric capsules and single but well-developed peristome are diagnostic. Confusion between this species and Anacolia laevisphaera may occur, especially with sterile material. In the latter, the apices of the distal leaves are often divergent when dry, spreading recurved when moist, with the inner basal laminal cells more differentiated than those of the basal angles compared with basal areolation in B. stricta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Bartramia halleriana is quite similar vegetatively to robust forms of B. pomiformis. However, the short, curved seta as long as or slightly longer than the capsule is distinctive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 103. FNA vol. 28, p. 102.
Parent taxa Bartramiaceae > Bartramia Bartramiaceae > Bartramia
Sibling taxa
B. brevifolia, B. halleriana, B. ithyphylla, B. pomiformis, B. potosica, B. subulata
B. brevifolia, B. ithyphylla, B. pomiformis, B. potosica, B. stricta, B. subulata
Name authority Bridel: Muscol. Recent. 2(3): 132, plate 1, fig. 5. (1803) Hedwig: Sp. Musc. Frond., 164. (1801)
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