Bartramia stricta
Bartramia
bartramia moss, rigid apple moss
bartramia moss
1–3 cm.
0.5–15 cm, erect, 2-fid, lacking subfloral whorl of branches; rounded in cross section, hyalodermis present, indistinct to distinct, epidermis not prorulose;
radiculose proximally, rhizoids papillose.
stiffly erect-appressed when dry, erect-spreading when moist, narrowly lanceolate, 2.5–4 mm;
base not sheathing, shoulders absent;
margins recurved from near base to distal acumen, serrulate distally, teeth single;
apex narrowly aristate;
costa excurrent, abaxially prominent, distal abaxial surface rough;
basal laminal cell walls thin;
medial and distal cells 12–20 × 4–6 µm, prorulae low.
not in distinct rows or rarely in 5 rows, erect-appressed to spreading or circinate, sometimes flexuose when dry, spreading when moist, narrowly lanceolate to linear, 2- or 3-stratose at margins or throughout;
base ± sheathing;
margins plane or revolute, entire proximally, serrate to serrulate distally, teeth single or paired;
apex acuminate or subulate;
costa subpercurrent to excurrent, abaxial surface prominent, rough, or sometimes obscure in distal limb;
basal laminal cells elongate to rectangular or linear;
distal cells subquadrate to oblong-linear, prorulose on both surfaces, walls firm.
0.1–1.5 cm, straight.
single (often 2 or 3 per perichaetium in B. halleriana), elongate or rarely short, straight or sometimes curved.
synoicous.
dioicous, autoicous, or synoicous;
perigonia gemmiform;
perichaetial leaves little differentiated from stem leaves (differentiated in B. ithyphylla).
erect, subglobose to ovoid, symmetric, 1.4–2 mm;
operculum conic convex;
peristome single;
exostome teeth 250–300 µm, striolate papillose proximally, smooth distally;
endostome absent.
inclined or sometimes erect, subglobose, globose, ovoid, or pyriform, furrowed when dry, mouth oblique;
annulus not distinct;
operculum convex to conic;
peristome double, single, or absent;
exostome teeth reddish brown, lanceolate, smooth or papillose, apically free;
endostome pale yellow, sometimes absent, segments keeled, cilia rudimentary or absent.
22–26 µm.
reniform to subspheric, coarsely to warty papillose.
asexual reproduction absent.
Bartramia stricta
Bartramia
Bartramia stricta is a widespread Old World species occurring infrequently in the southwestern portion of the flora area, with an isolated occurrence in western Canada. The strict, appressed leaves without shoulders and with relatively uniform areolation from base to acumen aid in recognition. When fertile, the erect, symmetric capsules and single but well-developed peristome are diagnostic. Confusion between this species and Anacolia laevisphaera may occur, especially with sterile material. In the latter, the apices of the distal leaves are often divergent when dry, spreading recurved when moist, with the inner basal laminal cells more differentiated than those of the basal angles compared with basal areolation in B. stricta.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species ca. 90 (7 in the flora).
Bartramia is generally recognized by its linear-lanceolate leaves and sheathing base with a distal lamina that is 2- or 3-stratose at the margins or throughout. The plants may be soft or rigid; the stem cortical cells have somewhat firm walls. The leaves are not plicate; the basal laminal cells are pale and smooth with walls thin or rarely thick toward the costa. The perichaetial leaves are usually less prorulose with more lax areolation; the exostome teeth are sometimes cleft distally and lack intermediate abaxial thickenings. Bartramia stricta lacks a sheathing leaf base but otherwise is in accord with the generic concept.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaves 4-7 mm; bases laxly or scarcely sheathing; stems to 10(-15) cm | → 2 |
1. Leaves usually 1.5-5 mm; bases sheathing (except B. stricta); stems to 5 cm | → 3 |
2. Setae short, equal to or slightly longer than capsule. | B. halleriana |
2. Setae many times longer than capsule. | B. pomiformis |
3. Leaf bases not sheathing, shoulders absent. | B. stricta |
3. Leaf bases sheathing, shoulders well developed | → 4 |
4. Costae obscure in distal limb; distal laminal cells 25-45 µm, prorulae low. | B. ithyphylla |
4. Costae prominent in distal limb; distal laminal cells 8-25 µm, prorulae high | → 5 |
5. Leaf shoulders delicate, often eroded; apices fragile, usually broken. | B. potosica |
5. Leaf shoulders firm, not eroded; apices usually intact | → 6 |
6. Basal laminal cell walls thick toward costa, thin toward margins; leaf margins strongly revolute. | B. brevifolia |
6. Basal laminal cell walls thin throughout; leaf margins plane distally. | B. subulata |
CalFlora
PNW Herbaria
WildflowerSearch
iNaturalist
USDA Plants Database
