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bartramia moss, straight-leaf apple-moss, straight-leaf bartramia

Haller's apple-moss, Haller's bartramia moss

Habit Plants in lax to dense tufts, soft green to glaucous. Plants in lax to dense tufts, greenish to yellowish brown.
Stems

1–3(–5) cm.

15 cm.

Leaves

stiffly erect when dry, spreading when moist, linear, 4–5 mm;

base sheathing, shoulders well developed, firm;

margins plane, serrulate to serrate distally, teeth paired distally;

apex acuminate, subulate;

costa excurrent, obscure in distal limb;

basal laminal cell walls thin;

medial and distal cells 25–45 × 5–7 µm, prorulae relatively low.

crisped when dry, erect-spreading or sometimes secund when moist, narrowly lanceolate to linear, 4–7 mm;

base laxly sheathing, shoulders well developed, firm;

margins revolute from shoulders to high in acumen, denticulate in shoulders, spinose-dentate distally, teeth paired;

apex long-subulate;

costa excurrent, prominent, distal abaxial surface rough;

basal laminal cell walls thin;

medial and distal cells 5–12 × 4(–10) µm, prorulae high.

Seta

0.8–3 cm, straight.

0.2(–0.8) cm, curved.

Sexual condition

synoicous;

perichaetial leaves somewhat longer than stem leaves, 6 mm, more strongly clasping.

autoicous or synoicous.

Capsule

inclined, subglobose to ovoid, asymmetric, 1 mm;

operculum short-conic;

peristome double;

exostome teeth 300–400 µm, strongly transversely barred, finely papillose proximally, smooth distally;

endostome basal membrane present, segments 1/2–2/3 length of teeth and somewhat adherent to them, smooth, cilia absent or rudimentary.

inclined, subglobose to pyriform, asymmetric, 1.5(–2.5) mm;

operculum conic convex;

peristome double;

exostome teeth 500–550 µm, granulose proximally, papillose distally;

endostome basal membrane high, segments slightly shorter than teeth, striate papillose, cilia rudimentary.

Spores

25–40 µm.

15–24 µm.

Bartramia ithyphylla

Bartramia halleriana

Phenology Capsules mature Jul–Nov. Spores 15-24 µm, mature Jun–Sep.
Habitat Soil, rock Crevices of shaded cliffs, rock outcrops in humid forests
Elevation low to high elevations (0-3800 m) (low to high elevations (0-12500 ft)) moderate elevations (200-500 m) (moderate elevations (700-1600 ft))
Distribution
from FNA
AK; CA; CO; ID; ME; MI; MT; NH; NV; OR; UT; WA; WY; AB; BC; NB; NL; NU; QC; YT; e Asia (Taiwan); n Africa; Greenland; Europe; s South America (Argentina); c Africa
[WildflowerSearch map]
from FNA
AB; BC; s South America; Europe; Asia; Pacific Islands (Hawaii, New Zealand); Australia (Tasmania)
Discussion

Bartramia ithyphylla is essentially an arctic-alpine species with disjunct populations in austral South America and the high mountains of Africa. In the flora area, the species frequents tundra and montane forest habitats with occasional occurrence at low to moderate elevations at northern latitudes. The glistening white leaf base is distinctive. The obscure costa in the distal limb and elongate distal laminal cells bearing low prorulae distinguish B. ithyphylla from other small species of the genus in the flora area. The distal leaves are sometimes divergent. Reports of Bartramia breviseta Lindberg [B. ithyphylla var. breviseta (Lindberg) Kindberg by some authors] from high elevations in Colorado likely represent misidentifications. In B. breviseta the capsules are overtopped by the perichaetial leaves (the seta is 1–3 mm), and the costa fills the acumen. As presently understood, B. breviseta is an arctic-alpine species of the Old World.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Bartramia halleriana is quite similar vegetatively to robust forms of B. pomiformis. However, the short, curved seta as long as or slightly longer than the capsule is distinctive.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 28, p. 104. FNA vol. 28, p. 102.
Parent taxa Bartramiaceae > Bartramia Bartramiaceae > Bartramia
Sibling taxa
B. brevifolia, B. halleriana, B. pomiformis, B. potosica, B. stricta, B. subulata
B. brevifolia, B. ithyphylla, B. pomiformis, B. potosica, B. stricta, B. subulata
Synonyms B. ithyphylla var. breviseta, B. ithyphylla subsp. rigidula
Name authority Bridel: Muscol. Recent. 2(3): 132, plate 1, fig. 6. (1803) Hedwig: Sp. Musc. Frond., 164. (1801)
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