Bartramia ithyphylla
Bartramia
bartramia moss, straight-leaf apple-moss, straight-leaf bartramia
bartramia moss
1–3(–5) cm.
0.5–15 cm, erect, 2-fid, lacking subfloral whorl of branches; rounded in cross section, hyalodermis present, indistinct to distinct, epidermis not prorulose;
radiculose proximally, rhizoids papillose.
stiffly erect when dry, spreading when moist, linear, 4–5 mm;
base sheathing, shoulders well developed, firm;
margins plane, serrulate to serrate distally, teeth paired distally;
apex acuminate, subulate;
costa excurrent, obscure in distal limb;
basal laminal cell walls thin;
medial and distal cells 25–45 × 5–7 µm, prorulae relatively low.
not in distinct rows or rarely in 5 rows, erect-appressed to spreading or circinate, sometimes flexuose when dry, spreading when moist, narrowly lanceolate to linear, 2- or 3-stratose at margins or throughout;
base ± sheathing;
margins plane or revolute, entire proximally, serrate to serrulate distally, teeth single or paired;
apex acuminate or subulate;
costa subpercurrent to excurrent, abaxial surface prominent, rough, or sometimes obscure in distal limb;
basal laminal cells elongate to rectangular or linear;
distal cells subquadrate to oblong-linear, prorulose on both surfaces, walls firm.
0.8–3 cm, straight.
single (often 2 or 3 per perichaetium in B. halleriana), elongate or rarely short, straight or sometimes curved.
synoicous;
perichaetial leaves somewhat longer than stem leaves, 6 mm, more strongly clasping.
dioicous, autoicous, or synoicous;
perigonia gemmiform;
perichaetial leaves little differentiated from stem leaves (differentiated in B. ithyphylla).
inclined, subglobose to ovoid, asymmetric, 1 mm;
operculum short-conic;
peristome double;
exostome teeth 300–400 µm, strongly transversely barred, finely papillose proximally, smooth distally;
endostome basal membrane present, segments 1/2–2/3 length of teeth and somewhat adherent to them, smooth, cilia absent or rudimentary.
inclined or sometimes erect, subglobose, globose, ovoid, or pyriform, furrowed when dry, mouth oblique;
annulus not distinct;
operculum convex to conic;
peristome double, single, or absent;
exostome teeth reddish brown, lanceolate, smooth or papillose, apically free;
endostome pale yellow, sometimes absent, segments keeled, cilia rudimentary or absent.
25–40 µm.
reniform to subspheric, coarsely to warty papillose.
asexual reproduction absent.
Bartramia ithyphylla
Bartramia
Bartramia ithyphylla is essentially an arctic-alpine species with disjunct populations in austral South America and the high mountains of Africa. In the flora area, the species frequents tundra and montane forest habitats with occasional occurrence at low to moderate elevations at northern latitudes. The glistening white leaf base is distinctive. The obscure costa in the distal limb and elongate distal laminal cells bearing low prorulae distinguish B. ithyphylla from other small species of the genus in the flora area. The distal leaves are sometimes divergent. Reports of Bartramia breviseta Lindberg [B. ithyphylla var. breviseta (Lindberg) Kindberg by some authors] from high elevations in Colorado likely represent misidentifications. In B. breviseta the capsules are overtopped by the perichaetial leaves (the seta is 1–3 mm), and the costa fills the acumen. As presently understood, B. breviseta is an arctic-alpine species of the Old World.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Species ca. 90 (7 in the flora).
Bartramia is generally recognized by its linear-lanceolate leaves and sheathing base with a distal lamina that is 2- or 3-stratose at the margins or throughout. The plants may be soft or rigid; the stem cortical cells have somewhat firm walls. The leaves are not plicate; the basal laminal cells are pale and smooth with walls thin or rarely thick toward the costa. The perichaetial leaves are usually less prorulose with more lax areolation; the exostome teeth are sometimes cleft distally and lack intermediate abaxial thickenings. Bartramia stricta lacks a sheathing leaf base but otherwise is in accord with the generic concept.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Leaves 4-7 mm; bases laxly or scarcely sheathing; stems to 10(-15) cm | → 2 |
1. Leaves usually 1.5-5 mm; bases sheathing (except B. stricta); stems to 5 cm | → 3 |
2. Setae short, equal to or slightly longer than capsule. | B. halleriana |
2. Setae many times longer than capsule. | B. pomiformis |
3. Leaf bases not sheathing, shoulders absent. | B. stricta |
3. Leaf bases sheathing, shoulders well developed | → 4 |
4. Costae obscure in distal limb; distal laminal cells 25-45 µm, prorulae low. | B. ithyphylla |
4. Costae prominent in distal limb; distal laminal cells 8-25 µm, prorulae high | → 5 |
5. Leaf shoulders delicate, often eroded; apices fragile, usually broken. | B. potosica |
5. Leaf shoulders firm, not eroded; apices usually intact | → 6 |
6. Basal laminal cell walls thick toward costa, thin toward margins; leaf margins strongly revolute. | B. brevifolia |
6. Basal laminal cell walls thin throughout; leaf margins plane distally. | B. subulata |
BC
CalFlora
Flora NW
MI Flora
PNW Herbaria
WildflowerSearch
iNaturalist
USDA Plants Database
