Balsamorhiza macrophylla |
Balsamorhiza careyana |
|
---|---|---|
cutleaf balsamroot, Idaho balsamroot, large-leaf balsamroot |
Carey's balsamroot |
|
Habit | Plants 30–45(–100) cm. | Plants (15–)20–60 cm. |
Basal leaves | green, ovate to lanceolate, 20–50+ × 8–15 cm (pinnatifid, lobes lanceolate, 20–80+ × 10–40 mm, entire or ± dentate), bases ± cuneate, ultimate margins usually entire (plane or weakly revolute, ciliate), apices obtuse to acute, faces scabrous or piloso-hirtellous to pilose (at least abaxial usually gland-dotted as well). |
blades green, rounded-deltate or deltate to triangular-deltate, 15–25 × 6–15 cm, bases cordate or hastate to truncate, margins usually entire, sometimes crenate (to dentate near bases), apices acute to attenuate, faces finely hispidulous to hirtellous (gland-dotted as well). |
Involucres | ± hemispheric, 20–30 mm diam. |
hemispheric to turbinate or campanulate, 12–20 mm diam. |
Ray laminae | 35–50+ mm. |
20–30(–40) mm (cypselae strigose or glabrous). |
Outer phyllaries | lance-ovate or lanceolate to lance-linear, 12–30(–40) mm, equaling or surpassing inner (margins ciliate), apices acute to attenuate. |
oblong to lanceolate or linear, 15–25 mm, usually surpassing inner, apices acute to attenuate. |
Heads | usually borne singly. |
usually (2–)3+, sometimes borne singly. |
2n | = 100 ± 2. |
= 38. |
Balsamorhiza macrophylla |
Balsamorhiza careyana |
|
Phenology | Flowering May–Jul. | Flowering (Mar–)May–Jun(–Jul). |
Habitat | Deep soils, rocky meadows, sagebrush scrublands, conifer forests | Dry scablands, semi-desert soils, openings in pine forests |
Elevation | 1000–2400 m (3300–7900 ft) | 500–1000 m (1600–3300 ft) |
Distribution |
ID; UT; WY
|
OR; WA
|
Discussion | Balsamorhiza macrophylla is a high polyploid; it occurs sympatrically with B. sagittata. It evidently arose from hybridization between B. sagittata and B. hispidula. Balsamorhiza macrophylla has the multi-branched caudices and massive taproots of the former, and the leaf dissection of the latter. No hybrids with other species are known. Presumably, the high-polyploid chromosome complement precludes interbreeding. Plants of var. idahoensis are smaller, are known only from southwestern Idaho and northeastern Utah, and differ from var. macrophylla by being pilose, with strongly shaggy-pilose involucres. More study may determine that var. idahoensis merits specific rank. The Utah populations are not well understood and deserve attention. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Hybrids involving Balsamorhiza careyana and B. deltoidea occur near the Columbia River Gorge; intermediates are found to the east and south. In northern Oregon, plants in some populations have glabrous cypselae and some populations are mixed. The hairiness may come from B. rosea; B. rosea and B. careyana may hybridize profusely, producing mostly plants with the stature of B. careyana and with the relatively short, brick-red ray corollas of B. rosea. Hybridization also occurs, occasionally, between B. careyana and B. sagittata; B. careyana also hybridizes with any species of sect. Balsamorhiza with which it comes in contact. Plants called Balsamorhiza careyana var. intermedia usually have crenate leaf margins and glabrous cypselae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 21, p. 98. | FNA vol. 21, p. 95. |
Parent taxa | Asteraceae > tribe Heliantheae > subtribe Ecliptinae > Balsamorhiza > subg. Balsamorhiza | Asteraceae > tribe Heliantheae > subtribe Ecliptinae > Balsamorhiza > subg. Artorhiza |
Sibling taxa | ||
Synonyms | B. hookeri var. idahoensis, B. macrophylla var. idahoensis | B. careyana var. intermedia |
Name authority | Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 350. (1840) | A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 81. (1849) |
Web links |