Baccharis pilularis |
Baccharis glutinosa |
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chaparral broom, coyote brush, dwarf chaparral false willow |
Douglas' falsewillow, marsh baccharis, salt marsh baccharis |
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Habit | Shrubs, 15–450 cm (prostrate and mat-forming to erect and rounded, much branched). | Perennials, 60–210 cm (rhizoma-tous, forming colonies). | ||||
Stems | spreading to ascending, dark brown, shiny, striate-angular, glabrous, often ± scurfy, usually resinous and sticky. |
erect to ascending, striate, glabrous, gland-dotted, resinous. |
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Leaves | present at flowering; sessile or short-petiolate; blades (1- or 3-nerved) oblanceolate to obovate, the smaller 5–40 × 2–15 mm (thick), bases cuneate, margins entire or coarsely dentate (teeth 3–9 distal to middles), faces glabrous, gland-dotted, resinous. |
present at flowering; short-petiolate; blades (1- or 3-nerved, larger prominently 3-nerved) lanceolate, 50–130 × 8–30 mm, bases tapered to petioles, margins entire or finely dentate, apices acute to acuminate, faces glabrous, black gland-dotted. |
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Involucres | hemispheric to campanulate; staminate 3.2–5 mm, pistillate 3–6 mm. |
hemispheric; staminate 3.5–5 mm, pistillate 3.8–4.8(–6) mm. |
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Pistillate florets | 19–43; corollas 2.5–3.5 mm. |
80–150; corollas 1.7–3 mm. |
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Staminate florets | 20–34, 3–4 mm. |
26–40; corollas 3.5–4 mm. |
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Phyllaries | ovate to lanceolate, 1–3 mm, margins yellowish, scarious, medians yellow proximally, green distally, apices obtuse to acute or acuminate (erose, abaxial faces papillose-scurfy). |
narrowly lanceolate, 2–4 mm, inner series ± equal, margins yellowish, medians green to purplish, apices acuminate, erose or ciliate, faces glabrous, gland-dotted, resinous. |
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Heads | (100–200+) in (leafy) paniculiform arrays. |
in dense, terminal, flat-topped, corymbiform arrays. |
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Cypselae | 1–2 mm, 8–10-nerved, glabrous; pappi 6–9 mm. |
0.6–1.5 mm, 5-nerved, glandular, hispidulous distally; pappi 2.6–4(–7) mm. |
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2n | = 18. |
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Baccharis pilularis |
Baccharis glutinosa |
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Phenology | Flowering Jul–Oct (all year). | |||||
Habitat | Moist salt marshes, coastal strands, stream edges, hillsides, railroads | |||||
Elevation | 0–1200 m (0–3900 ft) | |||||
Distribution |
CA; NM; OR; nw Mexico
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CA; OR; Mexico (Baja California)
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Discussion | Subspecies 2 (2 in the flora). Baccharis pilularis can be distinguished by its dark brown stems, and serrate, obovate to oblanceolate leaves. In addition, plants from some dunes of the California coast are prostrate, a growth form unique to this genus in our region. The common, weedy, widespread form is subsp. consanguinea, which is typically erect, with its larger leaves 15–40 mm. Subspecies pilularis is known only from exposed sandy dunes and bluffs along the central coast of California. Its growth habit is matlike, and its larger leaves are 5–15 mm. The prostrate habit of subsp. pilularis is strikingly different from the upright habit of subsp. consanguinea. C. B. Wolf (1935) demonstrated that in at least some populations, the distinction between prostrate and erect forms has a genetic basis. Transplants from the wild of the prostrate and erect forms retained their respective growth habits when grown together in a sheltered location and the morphology of seedlings reflected the habit of the parents. Wolf’s arguments for recognizing the forms as subspecies are further supported by the existence of prostrate cultivars in the horticultural trade. On the other hand, both erect and prostrate forms grow in proximity throughout the range of subsp. pilularis. In many areas the forms intergrade completely; in others they can be easily distinguished. Two subspecies are recognized here, notwithstanding difficulties in identifying habit from pressed specimens, or by observations of populations where both growth forms coexist. Further study is needed, perhaps utilizing molecular characters and detailed observations of native populations. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Occurring along the coast and in interior valleys, Baccharis glutinosa is recognized by the erect, simple stems growing in patches from rhizomes, large lanceolate leaves with three veins and blackish glandular dots, heads in dense, compact, terminal, corymbiform arrays and more or less uniform inner phyllaries. G. L. Nesom (1990h) noted that it is similar to forms of the South American species Baccharis pingraea de Candolle, and that the two taxa may be conspecific. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 29. | FNA vol. 20, p. 27. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | B. douglasii | |||||
Name authority | de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 407. (1836) | Persoon: Sym. Pl. 2: 425. (1807) | ||||
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