Atriplex tularensis
Atriplex mucronata
Bakersfield smallscale, Tulare orach, Tulare saltbush
quelite
aging red, terete or obscurely angled, brittle.
erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young.
alternate or proximalmost opposite;
blade lanceolate to ovate, 6–20 × 4–8 mm, base rounded, margin entire, apex acute to acuminate, gray scurfy.
alternate or proximalmost opposite or subopposite;
petiole to 1.5 cm or sessile;
blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above.
in small axillary dense glomerules, 4-merous.
solitary or in small axillary clusters, below or mixed with staminate.
fascicled in axils.
dark brown, 1–1.2 mm.
reddish brown, 2 mm.
bracteoles sessile, rhombic-ovate, acute or acuminate, 2.5–3.5 × 2.2–3 mm, united to middle, thin margin toothed, faces plane, scurfy.
bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged.
glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles.
= 18.
Atriplex tularensis
Atriplex mucronata
Of conservation concern.
H. M. Hall and F. E. Clements (1923) discussed the relationship of this plant to Atriplex cordulata, noting that, “A. tularensis is a much more slender plant, the leaves are narrower in proportion to their length,” and tapering to the base, never at all cordate. They noted further, that “The fruiting bracts are sometimes very similar in shape, but those of tularensis typically end in an abrupt acute tooth not present in cordulata and both bract and seed are always smaller.” Because of its small fruiting bracteoles it was placed within the Pusillae by P. C. Standley (1916). The erect stature, coupled with distinctive spreading-ascending and stiff (at least in dried material) leaves, and foliose bracts apparently distinguish A. tularensis from A. coronata.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment).
The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year.
H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well.
Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles.
Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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