Flora of North America species comparison

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Atriplex suberecta

Atriplex covillei

peregrine saltbush, sprawling saltbush

Coville's orach
Habit Herbs, annual or perennial, sprawling to ascending, 2–6 dm, branching from densely scaly base. Herbs, spreading, 1–4(–5) dm and as broad.
Stems

terete, sparsely scurfy when young.
Leaves

mostly alternate, shortly petiolate;

blade narrow to broadly rhomboid, lanceolate, oblanceolate, or elliptic, (8–)12–35(–42) × 6–16 mm, thin, margin coarsely and irregularly dentate, glabrescent adaxially, somewhat scurfy abaxially.

petiole to 1/2 as long as blade (becoming subsessile distally);

blade green or finally stramineous, (10–)20–50 × 6–30 mm, firm, base abruptly acute to narrowly cuneate, apex acute to attenuate, sparsely scurfy.
Staminate flowers

in subterminal, axillary glomerules.

in sessile glomerules in distal axils, often mixed with pistillate ones, staminate calyx deeply 5-cleft;

lobes obtuse, not appendaged.
Pistillate flowers

in axillary glomerules.

with calyx of (1–)3(–5) hyaline sepals.
Seeds

circular.

dark reddish brown, 1–1.5 mm.
Fruiting

bracteoles sessile or on the stipe to 0.5 mm, rhombic to obovate, almost flat to convex, 2.2–4 × 1.7–2.7 mm, thin or somewhat thickened in age, connate in basal 1/2, margin entire in basal 1/2, 2–4-toothed in distal 1/2, apex acute, scurfy.

bracteoles sessile or stipitate, 6–12 × 4–7 mm, margin mostly 3-lobed, with elongate terminal lobe triangular to lanceolate, 2 short rounded lobes at base or sides merely rounded at base, united to beyond middle.
2n

= 18.

Atriplex suberecta

Atriplex covillei
Phenology Flowering summer–fall. Flowering summer–fall.
Habitat Disturbed places, with other ruderal weeds Mixed saltbush-greasewood, rabbitbrush, warm desert shrub, and salt grass communities in saline substrates
Elevation 10-900 m (0-3000 ft) 800-1700 m (2600-5600 ft)
Distribution
from FNA
CA; UT; Australia; naturalized South Africa [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
Discussion

Endolepis covillei was treated within the synonymy of Atriplex phyllostegia (Torrey) S. Watson by H. M. Hall and F. E. Clements (1923).

H. C. Stutz et al. (1993) placed Atriplex covillei within Endolepis, based in large part on the presence of a perianth subtending the pistil within the fruiting bracteoles and on the lack of Kranz anatomy in the leaves. The pattern of venation is, nevertheless, very similar to that in species with Kranz anatomy. The presence of perianth scales in the pistillate flowers of A. covillei has been regarded as evidence of relationship with A. suckleyi. Despite placement of these taxa within Endolepis by Stutz et al. Atriplex covillei is possibly more closely allied to the morphologically similar and partially sympatric A. phyllostegia than it is to strongly dissimilar and the distantly disjunct A. suckleyi. Stutz and his associates placed great emphasis on the presence of reduced perianth segments subtending the pistil within the fruiting bracteoles of A. covillei. Calyces per se, otherwise known only in A. suckleyi and A. pleiantha, probably have arisen independently. Their presence does not necessarily indicate a close relationship. Stutz et al. pointed to other differences aside from the calyx of the pistillate flowers, and it is apparent that the two entities can stand as distinct species. To segregate A. covillei within a separate genus and to ally it with a species to which its relationships are obscure at best, stretches logic beyond reason.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 4, p. 343. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Atriplex > sect. Semibaccata Chenopodiaceae > Atriplex > subg. Obione > sect. Covilleiae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Synonyms Endolepis covillei
Name authority I. Verdoorn: Bothalia 6: 418, figs. 2, 3(2). (1954) (Standley) J. F. Macbride: Contr. Gray Herb. 53: 11. (1918)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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