Atriplex serenana
Atriplex suckleyi
bracteate orach, bractscale, saltscale, stinking orach
Suckley's orach
terete, often tinged with red, sparsely mealy or glabrous.
many, subsessile or very short petiolate;
blade subconcolorous, lanceolate to oblong, elliptic, or oval, (8–)10–30(–40) × 3–12(–15) mm, margin sharply dentate to entire.
numerous, alternate, sessile;
blade lanceolate to elliptic or less commonly ovate, 7–35 × (2–)4–10(–11) mm, thick and succulent, base acute, margin entire, apex acute or acuminate, glaucous, sparsely farinose when young.
in glomerules in terminal spikes or panicles 3–20 cm, or reduced to solitary, rounded, terminal glomerule.
in small glomerules, these in distal axils or in short, dense or interrupted, mostly simple, terminal spikes, perianth cup-shaped, frequently pinkish, lobes each with fleshy crest.
in small clusters, axillary.
solitary or few in leaf axils, calyx within bracteoles of 3 or 4, entire or lobed, distinct, obtuse, hyaline sepals.
brown, 1–1.3(–1.5) mm.
brown, 1.5 × 1.2 mm.
bracteoles sessile or subsessile (stipe to 1 mm), cuneate-orbicular to obovate, somewhat compressed, 2.1–3.5 × (1.7–)2–3.7 mm, united 1/2 of length, margin sharply and often slenderly toothed beyond middle, faces often rather strongly veined, smooth or with 1 or more slender or flattened appendages.
bracteoles small and difficult to find, sessile, ovate, 2 × 1.5 mm, membranous, united to apex, without appendages, scurfy, each pistil subtended by a perianth.
= 18.
Atriplex serenana
Atriplex suckleyi
Varieties 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
At specific rank, the epithet suckleyi clearly has priority. Stutz et al. (1993) resurrected Endolepis Torrey and placed within it two completely disparate species, E. dioica and E. covillei Standley [Atriplex covillei (Standley) J. F. Macbride]. The interpretation of the genus by Stutz et al. stands on the presence of perianth segments subtending the ovary within the fruiting bracteoles, lack of Kranz leaf anatomy, and other more equivocal characteristics. The shared features hardly indicate near affinity, however. The two taxa are otherwise grossly dissimilar. Fundamentally, the genus Endolepis as resurrected by H. C. Stutz et al. stands on the basis of a single character: the presence of perianth segments. Kranz anatomy rises and falls, both within subg. Atriplex and subg. Obione. Thus, coincidence of the non-Kranz criterion is subject to interpretation. Perianth segments subtending the ovary within the enclosing bracteoles, per se, appear to be of independent origin. And, the peculiar nature of the staminate calyx in A. suckleyi (a major determining condition in establishment of the genus Endolepis by Torrey) is not present in A. covillei. Certainly the two species included within Endolepis appear to be as closely allied to other species of Atriplex as they are to each other.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Staminate inflorescence an elongate panicle or spike bearing numerous, beadlike glomerules; fruiting bracteoles ± 1-veined; mainly of central valley and Owens Valley, to w Nevada, also coastal and insu- lar, s California | var. serenana |
1. Staminate inflorescence of solitary, terminal, subglobose glomerule; fruiting bracteoles 3-veined; coastal s California | var. davidsonii |
CA
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