Atriplex semibaccata |
Atriplex suckleyi |
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Australian saltbush, berry saltbush, creeping saltbush |
Suckley's orach |
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Habit | Herbs or subshrubs, perennial, decumbent-prostrate, unarmed, mainly 0.5–8 dm and spreading to 15+ dm wide, unarmed, white scurfy when young; branches not angled. | Herbs, spreading, branching from base, (0.3–)0.5–3(–4) dm and as broad. |
Stems | terete, often tinged with red, sparsely mealy or glabrous. |
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Leaves | many, alternate, subsessile or short petiolate; blade 1-veined, spatulate or obovate to oblong or elliptic, mainly 5–30(–40) × 2–9(–12) mm, base attenuate, margin remotely dentate to subentire, apex obtuse. |
numerous, alternate, sessile; blade lanceolate to elliptic or less commonly ovate, 7–35 × (2–)4–10(–11) mm, thick and succulent, base acute, margin entire, apex acute or acuminate, glaucous, sparsely farinose when young. |
Staminate flowers | in small, terminal, leaf-bracteate glomerules 1.5 mm wide. |
in small glomerules, these in distal axils or in short, dense or interrupted, mostly simple, terminal spikes, perianth cup-shaped, frequently pinkish, lobes each with fleshy crest. |
Pistillate flowers | solitary or in few-flowered clusters in almost all but distalmost leaves. |
solitary or few in leaf axils, calyx within bracteoles of 3 or 4, entire or lobed, distinct, obtuse, hyaline sepals. |
Seeds | dimorphic: black, 1.5–1.7 mm, or brown, 2 mm. |
brown, 1.5 × 1.2 mm. |
Fruiting | bracteoles red-fleshy at maturity, sessile or short stipitate, strongly veined, rhombic, convex, 3–6.6 × 2.8–4.5 mm, united at base, margin toothed, apex obtuse to acute. |
bracteoles small and difficult to find, sessile, ovate, 2 × 1.5 mm, membranous, united to apex, without appendages, scurfy, each pistil subtended by a perianth. |
2n | = 18. |
|
Atriplex semibaccata |
Atriplex suckleyi |
|
Phenology | Flowering spring–early winter. | Flowering late summer–fall. |
Habitat | Saline waste places, along roads and sidewalks, in marshes, in various plant communities | Alkaline or saline, typically fine-textured substrates, often on shale or clay barrens, sometimes with other Atriplex spp., sagebrush, and grasses |
Elevation | 10-1000 m (0-3300 ft) | (400-)1200-2200 m ((1300-)3900-7200 ft) |
Distribution |
AZ; CA; DC; NM; NV; TX; UT; WA; Australia [Introduced in North America]
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CO; MT; ND; SD; WY; AB; SK |
Discussion | The red-fleshy fruiting bracteoles are diagnostic of this introduced perennial, which is multi-stemmed from an often buried woody caudex. The Australian species Atriplex muelleri Bentham is somewhat similar. It has been has reported, but not verified, in the North American flora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
At specific rank, the epithet suckleyi clearly has priority. Stutz et al. (1993) resurrected Endolepis Torrey and placed within it two completely disparate species, E. dioica and E. covillei Standley [Atriplex covillei (Standley) J. F. Macbride]. The interpretation of the genus by Stutz et al. stands on the presence of perianth segments subtending the ovary within the fruiting bracteoles, lack of Kranz leaf anatomy, and other more equivocal characteristics. The shared features hardly indicate near affinity, however. The two taxa are otherwise grossly dissimilar. Fundamentally, the genus Endolepis as resurrected by H. C. Stutz et al. stands on the basis of a single character: the presence of perianth segments. Kranz anatomy rises and falls, both within subg. Atriplex and subg. Obione. Thus, coincidence of the non-Kranz criterion is subject to interpretation. Perianth segments subtending the ovary within the enclosing bracteoles, per se, appear to be of independent origin. And, the peculiar nature of the staminate calyx in A. suckleyi (a major determining condition in establishment of the genus Endolepis by Torrey) is not present in A. covillei. Certainly the two species included within Endolepis appear to be as closely allied to other species of Atriplex as they are to each other. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 343. | FNA vol. 4, p. 345. |
Parent taxa | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Semibaccata | Chenopodiaceae > Atriplex > subg. Obione > sect. Endolepis |
Sibling taxa | ||
Synonyms | A. flagellaris | Endolepis suckleyi, A. endolepis, A. ovata, Endolepis dioica, Endolepis ovata, Kochia dioica, Salsola dioica |
Name authority | R. Brown: Prodr., 406. (1810) | (Torrey) Rydberg: Mem. New York Bot. Gard. 1: 134. (1900) |
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