Atriplex semibaccata |
Atriplex californica |
|
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Australian saltbush, berry saltbush, creeping saltbush |
California orach, California saltbush |
|
Habit | Herbs or subshrubs, perennial, decumbent-prostrate, unarmed, mainly 0.5–8 dm and spreading to 15+ dm wide, unarmed, white scurfy when young; branches not angled. | Herbs, monoecious or dioecious, prostrate or procumbent-decumbent, from fleshy fusiform or variously shaped taproot. |
Stems | many branched, subterete, 1.5–5 dm, white scurfy when young. |
|
Leaves | many, alternate, subsessile or short petiolate; blade 1-veined, spatulate or obovate to oblong or elliptic, mainly 5–30(–40) × 2–9(–12) mm, base attenuate, margin remotely dentate to subentire, apex obtuse. |
numerous, often crowded, alternate or proximalmost opposite; blade narrowly lanceolate to narrowly oblanceolate or elliptic, 4–20 × 1–5 mm, acute at both ends, gray scurfy. |
Staminate flowers | in small, terminal, leaf-bracteate glomerules 1.5 mm wide. |
in terminal bracteate spikes, or mixed with pistillate in rather dense axillary clusters, 4-merous. |
Pistillate flowers | solitary or in few-flowered clusters in almost all but distalmost leaves. |
|
Seeds | dimorphic: black, 1.5–1.7 mm, or brown, 2 mm. |
dark (black), 1–2 mm. |
Fruiting | bracteoles red-fleshy at maturity, sessile or short stipitate, strongly veined, rhombic, convex, 3–6.6 × 2.8–4.5 mm, united at base, margin toothed, apex obtuse to acute. |
bracteoles sessile, rhombic-ovate to ovate, scarcely united, 2.5–4 mm, margin entire, acute. |
2n | = 18. |
|
Atriplex semibaccata |
Atriplex californica |
|
Phenology | Flowering spring–early winter. | Flowering Apr–Nov. |
Habitat | Saline waste places, along roads and sidewalks, in marshes, in various plant communities | Sea bluffs, sandy coasts, crevices in sea cliffs, coastal strands, edges of coastal salt marsh, coastal sage scrub |
Elevation | 10-1000 m (0-3300 ft) | 0-50 m (0-200 ft) |
Distribution |
AZ; CA; DC; NM; NV; TX; UT; WA; Australia [Introduced in North America]
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CA; Mexico
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Discussion | The red-fleshy fruiting bracteoles are diagnostic of this introduced perennial, which is multi-stemmed from an often buried woody caudex. The Australian species Atriplex muelleri Bentham is somewhat similar. It has been has reported, but not verified, in the North American flora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. M. Hall and F. E. Clements (1923) placed great emphasis on the inferior radicle, dioecious habit, and free bracts in stating that there are no close relatives of Atriplex californica in North America. P. C. Standley (1916) regarded the radicle as lateral or superior, not inferior, and placed the species at the beginning of his treatment of the American species. Plants of A. californica, however, form a mirror-image, matched pair with A. watsonii, with which they share habit, leaf conformation, staminate glomerules arranged, at least partially, in terminal interrupted spikes, and Kranz anatomy, but from which they differ in the radicle position, valves of the fruiting bracteoles free beyond the middle, monoecious habit, and mostly alternate leaves. The interpretation by Hall and Clements of radicle position as fundamental is made problematic by the apparent random occurrence of a great many morphologic features from place to place within the genus and often within the taxon. Each character fails individually as having definitive taxonomic importance, making difficult or impossible an ultimately satisfactory phylogenetic arrangement. It is not, however, the character that makes the species, rather, it is the entire syndrome of features that constitutes the taxon. Most certainly A. californica is more closely allied to American taxa than to those from other regions of the world. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 343. | FNA vol. 4, p. 366. |
Parent taxa | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Semibaccata | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Californicae |
Sibling taxa | ||
Synonyms | A. flagellaris | |
Name authority | R. Brown: Prodr., 406. (1810) | Moquin-Tandon: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(2): 98. (1849) |
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