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sack saltbush, sack saltweed, stalk orach, stalk orache

Kleberg orach, Kleberg's saltbush

Habit Herbs, erect, forming rounded clumps. Herbs, with ligneous vertical taproot 5–9 mm thick; bark pale.
Stems

mainly 0.5–4(–5) dm;

stems usually branched from base, terete, stout, angled;

herbage scurfy.

erect, diffuse;

branches alternate, numerous, horizontal or distally ascending, terete, 1.5–4 dm, densely white farinose when young, glabrate in age;

bark becoming pale brownish white and flaky;

internodes mostly shorter than 1(–2.4) cm.

Leaves

alternate or proximalmost subopposite, short petiolate or distal ones sessile;

petiole 2–5 mm;

blade cordate-ovate or subreniform to ovate to deltoid-ovate or oval, mainly 6–40 × 4–30 mm, base truncate to subcordate or broadly cuneate, margin entire or in some subhastately lobed or sometimes undulate-dentate, apex acute to rounded.

alternate, proximalmost subopposite, sessile;

blade ovate-deltoid, 0.5–1.5(–2.5) × 5–12(–15) mm, rather firm and flat, base rounded, truncate, or slightly clasping, margin entire or toothed, apex acute, very densely canescent-farinose with a greenish yellow tinge.

Flowers

sessile, axillary, inconspicuous, mostly in leafy lateral branches with extremely short internodes, arising toward tips of secondary branches.

Staminate flowers

in most distal axils, 2 mm wide;

sepals 3–5, mostly hyaline, curved elliptic, 1.5–1.7 mm, mucronulate, farinose dorsally.

Pistillate flowers

usually in fascicles of 1–3 in proximal axils.

densely farinose;

bracteoles adnate to ovary.

Seeds

brownish to whitish, 1.5–2.3 mm.

dark reddish brown, round-lenticular, 1.5 mm wide, shining;

radicle superior.

Staminate

glomerules in distal axils or in short, naked, terminal (early deciduous) panicles;

flowers with 5-parted calyx.

Fruiting

bracteoles monomorphic or dimorphic, larger ones on stipes (2–)4–8(–15) mm, others sessile, united at base, round-triangular or suborbicular, 4–6 mm, irregularly and coarsely dentate and with ± densely beset with flat, cristate, or hornlike appendages, smaller bracteoles (lacking or rare in var. asterocarpa) in same axils, oblong to cuneate, 3–4 mm, apex truncate, dentate only at summit, faces smooth.

bracteoles variably and irregularly 3–7-cleft, ovate-orbicular, 3.1–4.7 × (3.2–)4–7 mm, typically somewhat constricted below middle, with terminal lobes (1–)1.5–2.8 mm, densely scurfy, faces doubly cristate or smooth.

2n

= 18.

Atriplex saccaria

Atriplex klebergorum

Phenology Flowering summer.
Habitat In silty or clay loam soils
Distribution
from FNA
AZ; CO; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; of conservation concern; near sea level
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The species was noted by its author as apparently belonging in sect. Argenteae Standley; I concur with that alignment. The overall shape of the bracteoles with a subterminal constriction is reminiscent of those of Atriplex powellii, but the bracteole shape is otherwise distinctive, and the long marginal teeth and occasional elongate cristate processes on the faces are unmatched elsewhere in the Argenteae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves mainly ovate to cordate in profile, the distalmost bracteate ones (and sometimes all) often broadly so in age; fruiting bracteoles with stipes to 10 mm, but seldom exceeding 6 mm; Arizona, New Mexico, Wyoming, s to Texas
var. saccaria
1. Leaves oval to rhombic, the bases truncate to cuneate-attenuate; fruiting bracteoles with stipes of various length; sw Utah and adjacent states
→ 2
2. Leaves oval to suborbiculate or less commonly elliptic; fruiting bracteoles with hornlike appendages, borne on stipes mainly 2-6 mm
var. asterocarpa
2. Leaves rhombic to oval or triangular, base cuneate-attenuate; fruiting bracteoles with flattened processes, borne on stipes mainly 2-15 mm (sessile in some)
var. cornuta
Source FNA vol. 4. FNA vol. 4, p. 353.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Saccariae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Argenteae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. saccaria var. asterocarpa, A. saccaria var. cornuta, A. saccaria var. saccaria
Synonyms A. truncata var. saccaria, Obione saccaria
Name authority S. Watson: Proc. Amer. Acad. Arts 9: 112. (1874) M. C. Johnston: SouthW. Naturalist 6: 49. (1961)
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