Atriplex pentandra |
Atriplex tatarica |
|
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seas hore orach |
Tatarian orach, Tatarian orache |
|
Habit | Herbs, annual or perennial, sprawling to erect, often suffrutescent at the base, much branched and clump-forming, 3–10 dm. | Herbs, much branched, forming tangled or spreading masses. |
Stems | obtusely angled, finely scurfy when young. |
with branches divaricate or ascending, terete or obtusely angled, 2–10(–15) dm, sparsely scurfy when young. |
Leaves | sessile or short petiolate; blade paler abaxially, oblong or rhombic-ovate to broadly obovate or narrowly oblong or narrowly elliptic, 10–30 × (1–)3–15 mm, thin, base rounded to cuneate, margin repand-dentate or sinuate-dentate to undulate or distal ones or all of them entire, apex rounded to acute, mucronate, densely white scurfy abaxially, grayish green and usually glabrate adaxially. |
alternate (or the proximalmost opposite), long petiolate becoming nearly sessile distally, blade ovate to triangular, 15–50(–60) × 10–40 mm, base subhastate or cuneate, margin deeply or shallowly sinuate-dentate with acute or obtuse teeth, apex acute or obtuse, distalmost bracteate blades becoming entire and linear or oblong-linear. |
Staminate flowers | in short, dense, naked terminal spikes or panicles; calyx 5-cleft, lobes green keeled. |
in glomerules borne in slender, naked or sparingly bracteate (at base), mostly interrupted simple or paniculate spikes, calyx 5-cleft. |
Pistillate flowers | fascicled in axils. |
fascicled in distal axils. |
Seeds | brown, 1–1.5 mm. |
brown, 1.5–2 mm; radicle inferior, ascending. |
Fruiting | bracteoles sessile or with stipes to 0.5 mm, broadly cuneate-orbiculate, compressed, (2–) 2.5–4.5 × (1.5–)2.6–5 mm, usually as broad as or broader than long, much thickened at maturity, united only at truncate or broadly cuneate base, margin deeply and acutely dentate, faces with 2, sometimes swollen, dentate crests or covered with irregular, conic-acute, corky tubercles, seldom smooth. |
bracteoles strongly 3-veined and reticulate, sessile or short stipitate, ovate-rhombic or subflabelliform, 4–8 × 3–7 mm, moderately compressed, united from narrowed base to middle, margin broad, foliaceous, coarsely dentate, indurate at maturity, faces tuberculate or smooth. |
Atriplex pentandra |
Atriplex tatarica |
|
Phenology | Flowering summer–fall. | Flowering late summer–fall. |
Habitat | Sandy seashores, coastal salt marshes | Atlantic and Gulf coasts, ballast and waste grounds |
Elevation | 0-50 m (0-200 ft) | 0-50 m (0-200 ft) |
Distribution |
AL; CT; FL; GA; LA; MA; MS; NC; SC; TX; West Indies; South America (Venezuela and Colombia to Peru) |
AL; CT; FL; MA; NH; NJ; PA; Europe [Introduced in North America] |
Discussion | D. S. Correll and M. C. Johnston (1970), treated both Atriplex texana and A. wardii at the specific level. The diagnostic features used to distinguish them, dentate versus entire leaf blades and smooth versus tuberculate faces of fruiting bracteoles, fail singly and in combination. Both taxa were regarded by H. M. Hall and F. E. Clements (1923) as variants of typical A. pentandra, and this worker tentatively agrees with those authors’ conclusions. Specimens from Galveston, Texas—the type locality of A. wardii—are apparently intermediate with A. mucronata (see below) and form the basis on which A. wardii was founded. Some of those specimens have been annotated by me and others as A. mucronata. The plants have entire leaves more closely matching those of A. mucronata, but the small fruiting bracteoles, though usually lacking tubercles on the faces, are of similar proportions to those of typical A. pentandra. Atriplex pentandra and A. mucronata are certainly very closely allied. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
H. M. Hall and F. E. Clements (1923) treated Atriplex tatarica at specific level with the above distribution (as an introduced ballast plant of rare occurrence), but did not recognize the closely similar A. laciniata, which is currently the species recognized for coastal eastern America. They evidently interpreted A. tatarica to include A. lampa Gillies as identified by P. C. Standley (1916). Atriplex lampa is a shrubby species from South America and does not figure in consideration of North American taxa. Atriplex tatarica includes a complex of forms and varieties in Flora URSS (M. M. Iljin 1936). In Flora Europaea (P. Aellen 1964b) A. tatarica and A. laciniata are separated in the key by the glomerules of A. tatarica being borne in terminal, leafless, often long panicles, whereas those of A. laciniata are borne axillary or in leafy clusters. The species differ otherwise in stature, A. laciniata being a dwarf plant up to 30 cm and with small leaves, A. tatarica being a robust annual up to 150 cm, and with large leaves. The illustration by Hall and Clements (fig. 38) shows a plant with leafy panicle. Atriplex tatarica has been ignored by H. A. Gleason and A. Cronquist (1991) and by all other authors of floras covering the northeastern United States. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 362. | FNA vol. 4, p. 341. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Axyris pentandra, A. texana, A. wardii | |
Name authority | (Jacquin) Standley: in N. L. Britton et al., N. Amer. Fl. 21: 54. (1916) | Linnaeus: Sp. Pl. 2: 1053. (1753) |
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