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seas hore orach

Parish's brittlescale, Parish's saltbush, Parrish's brittlescale

Habit Herbs, annual or perennial, sprawling to erect, often suffrutescent at the base, much branched and clump-forming, 3–10 dm. Herbs, erect or spreading to prostrate, 0.5–3 dm; branches almost horizontal to ascending, fragile, white scurfy or villous (in var. parishii).
Stems

obtusely angled, finely scurfy when young.

Leaves

sessile or short petiolate;

blade paler abaxially, oblong or rhombic-ovate to broadly obovate or narrowly oblong or narrowly elliptic, 10–30 × (1–)3–15 mm, thin, base rounded to cuneate, margin repand-dentate or sinuate-dentate to undulate or distal ones or all of them entire, apex rounded to acute, mucronate, densely white scurfy abaxially, grayish green and usually glabrate adaxially.

numerous, all or nearly all opposite or almost all alternate, distal ones imbricate or widely separated, tending to recurve;

blade lanceolate to ovate, (2–)4–10 × 3–8 mm, rigid, base mostly rounded to cordate, margin entire, gray to white, densely scurfy (or hairy).

Staminate flowers

in short, dense, naked terminal spikes or panicles;

calyx 5-cleft, lobes green keeled.

mostly in distal axils pistillate in proximal axils, or mostly in terminal spike (var. persistens), or partly so (var. subtilis).

Pistillate flowers

fascicled in axils.

Seeds

brown, 1–1.5 mm.

dark brown or almost black, 0.8–1.5 mm.

Fruiting

bracteoles sessile or with stipes to 0.5 mm, broadly cuneate-orbiculate, compressed, (2–) 2.5–4.5 × (1.5–)2.6–5 mm, usually as broad as or broader than long, much thickened at maturity, united only at truncate or broadly cuneate base, margin deeply and acutely dentate, faces with 2, sometimes swollen, dentate crests or covered with irregular, conic-acute, corky tubercles, seldom smooth.

bracteoles sessile, ovate or rhombic, slightly compressed to thickened, 2–3.5(–4) mm and about as broad or sometimes broader, often subhastately lobed, united 1/2 of length, entire or with few teeth on each side, tuberculate on 1 or both faces.

Atriplex pentandra

Atriplex parishii

Phenology Flowering summer–fall.
Habitat Sandy seashores, coastal salt marshes
Elevation 0-50 m (0-200 ft)
Distribution
from FNA
AL; CT; FL; GA; LA; MA; MS; NC; SC; TX; West Indies; South America (Venezuela and Colombia to Peru)
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

D. S. Correll and M. C. Johnston (1970), treated both Atriplex texana and A. wardii at the specific level. The diagnostic features used to distinguish them, dentate versus entire leaf blades and smooth versus tuberculate faces of fruiting bracteoles, fail singly and in combination. Both taxa were regarded by H. M. Hall and F. E. Clements (1923) as variants of typical A. pentandra, and this worker tentatively agrees with those authors’ conclusions. Specimens from Galveston, Texas—the type locality of A. wardii—are apparently intermediate with A. mucronata (see below) and form the basis on which A. wardii was founded. Some of those specimens have been annotated by me and others as A. mucronata. The plants have entire leaves more closely matching those of A. mucronata, but the small fruiting bracteoles, though usually lacking tubercles on the faces, are of similar proportions to those of typical A. pentandra. Atriplex pentandra and A. mucronata are certainly very closely allied.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (5 in the flora).

The Atriplex parishii complex consists of a series of microphyllous, low clump-forming annuals apparently disjunct from each other in the Central Valley of California and in near coastal southern California. Often they occupy vernal pools that dry as the season progresses; the substrates in all cases evidently are saline or alkaline, or both. For the most part, the bracteate distal leaves are cordate to rounded at the base, and spreading to spreading-ascending, and the fruiting bracteoles are mainly less than 3.5 mm in length.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruiting bracteoles persistent, ± thickened and/or tuberculate on 1 or both faces
→ 2
1. Fruiting bracteoles deciduous, compressed, faces smooth or tuberculate
→ 3
2. Stems, even in inflorescences, mainly over 1.5 mm thick; staminate flowers borne mainly in terminal spikes; fruiting bracteoles 2.5-3.5(-4) mm; Colusa, Glenn, Merced, Solano, Stanislaus, and Tulare counties.
var. persistens
2. Stems all or at least above, filiform, less than 1.5 mm thick; staminate flowers borne mainly in axillary glomerules, only sometimes somewhat spicate; fruiting bracteoles 2.4-2.8 mm; Fresno, Kings, Kern, Madera, and Tulare counties
var. subtilis
3. Plants with elongate slender hairs in inflorescences; foliose bracts of inflorescence spreading, tips ± recurved; plants of Los Angeles (or Orange), Riverside, and San Diego counties
var. parishii
3. Plants merely scurfy in inflorescences; foliose bracts of inflorescence spreading-ascending, tips not or not especially recurved
→ 4
4. Leaves all or mainly opposite, 3-7 mm; fruiting bracteoles with appendages not in rows, on abaxial surface only; Glenn, Solano, and Yolo counties
var. depressa
4. Leaves all or mainly alternate, 4-10 mm; fruiting bracteoles with faces smooth; Tulare and Kern counties
var. minuscula
Source FNA vol. 4, p. 362. FNA vol. 4, p. 356.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. parishii var. depressa, A. parishii var. minuscula, A. parishii var. parishii, A. parishii var. persistens, A. parishii var. subtilis
Synonyms Axyris pentandra, A. texana, A. wardii Obione parishii
Name authority (Jacquin) Standley: in N. L. Britton et al., N. Amer. Fl. 21: 54. (1916) S. Watson: Proc. Amer. Acad. Arts 17: 377. (1882)
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