Atriplex pentandra |
Atriplex littoralis |
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seas hore orach |
grassleaf orache, linear-leaf orache, narrow-leaf atriplex, narrow-leaf orache |
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Habit | Herbs, annual or perennial, sprawling to erect, often suffrutescent at the base, much branched and clump-forming, 3–10 dm. | Herbs, monoecious or subdioecious, 2.5–7.5(–10) dm. |
Stems | obtusely angled, finely scurfy when young. |
erect or forming sprawling tangled clumps and mostly green; branches erect-ascending, proximal ones opposite and ascending, sparsely scurfy when young, obtusely angled. |
Leaves | sessile or short petiolate; blade paler abaxially, oblong or rhombic-ovate to broadly obovate or narrowly oblong or narrowly elliptic, 10–30 × (1–)3–15 mm, thin, base rounded to cuneate, margin repand-dentate or sinuate-dentate to undulate or distal ones or all of them entire, apex rounded to acute, mucronate, densely white scurfy abaxially, grayish green and usually glabrate adaxially. |
alternate or only proximalmost opposite, all shortly petiolate; blade green on both surfaces, linear, linear-oblong, linear-lanceolate, or oblong, (10–)25–80(–120) × 2–8(–12) mm, thin, gradually narrowed, margin mostly entire or some sinuate-dentate with antrorse teeth, apex obtuse to acuminate. |
Inflorescences | of long, dense or interrupted hairy spikes often forming panicle to 20 cm; staminate flowers 4–5-merous. |
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Staminate flowers | in short, dense, naked terminal spikes or panicles; calyx 5-cleft, lobes green keeled. |
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Pistillate flowers | fascicled in axils. |
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Seeds | brown, 1–1.5 mm. |
dimorphic: brown, 2–2.8 mm wide, round and ± flattened, or black, 1.5–2 mm wide, round, evenly convex; radicle inferior. |
Fruiting | bracteoles sessile or with stipes to 0.5 mm, broadly cuneate-orbiculate, compressed, (2–) 2.5–4.5 × (1.5–)2.6–5 mm, usually as broad as or broader than long, much thickened at maturity, united only at truncate or broadly cuneate base, margin deeply and acutely dentate, faces with 2, sometimes swollen, dentate crests or covered with irregular, conic-acute, corky tubercles, seldom smooth. |
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Pistillate | bracteoles sessile, triangular to ovate or ovate-rhombic, (3–)5–7 mm, mostly denticulate, rarely subentire, faces tuberculate almost distinct, green becoming brown or black. |
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2n | = 18. |
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Atriplex pentandra |
Atriplex littoralis |
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Phenology | Flowering summer–fall. | Flowering summer. |
Habitat | Sandy seashores, coastal salt marshes | Sea beaches and other saline habitats, old ports and ballast dumps |
Elevation | 0-50 m (0-200 ft) | mainly below 100 m (mainly below 300 ft) |
Distribution |
AL; CT; FL; GA; LA; MA; MS; NC; SC; TX; West Indies; South America (Venezuela and Colombia to Peru) |
IL; IN; MA; ME; MI; NH; OH; PA; NB; NS; PE; Europe [Introduced in North America] |
Discussion | D. S. Correll and M. C. Johnston (1970), treated both Atriplex texana and A. wardii at the specific level. The diagnostic features used to distinguish them, dentate versus entire leaf blades and smooth versus tuberculate faces of fruiting bracteoles, fail singly and in combination. Both taxa were regarded by H. M. Hall and F. E. Clements (1923) as variants of typical A. pentandra, and this worker tentatively agrees with those authors’ conclusions. Specimens from Galveston, Texas—the type locality of A. wardii—are apparently intermediate with A. mucronata (see below) and form the basis on which A. wardii was founded. Some of those specimens have been annotated by me and others as A. mucronata. The plants have entire leaves more closely matching those of A. mucronata, but the small fruiting bracteoles, though usually lacking tubercles on the faces, are of similar proportions to those of typical A. pentandra. Atriplex pentandra and A. mucronata are certainly very closely allied. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4, p. 362. | FNA vol. 4. |
Parent taxa | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis |
Sibling taxa | ||
Synonyms | Axyris pentandra, A. texana, A. wardii | A. hastata var. littoralis, A. patula var. littoralis, A. patula subsp. littoralis |
Name authority | (Jacquin) Standley: in N. L. Britton et al., N. Amer. Fl. 21: 54. (1916) | Linnaeus: Sp. Pl. 2: 1054. (1753) |
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